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. 2025 Jan 1;15(3):1164–1184. doi: 10.7150/thno.104529

Table 2.

Functions of PSPs in TME

Related properties Enzyme Type of cell Mechanism/pathway Ref.
Hypoxic adaption PP1 Pancreatic cancer PPP1R1B promotes P53 degradation 16
PP2A CRC PP2A-B55α protects HIF1A from hydroxylation and degradation 92
Glioblastoma multiforme PP2A mediates cell cycle inhibition and reduced ATP consumption 103
Calcineurin Prostate cancer Calcineurin facilitates accumulation of HIF1A 93
PP5 Various tumor cells PP5 suppresses ASK-1/MKK4/JNK pathway 104
PPM1B Kidney cancer PPM1B regulates TBK1 signaling 105
PPM1G Renal adenocarcinoma PPM1G negatively regulates HIF1A expression 94
PHLPP Colon cancer PHLPP is negatively regulated by HIF1A 96
Metabolic reprogramming PP1 Gastric cancer PP1 regulates AKT/HIF1A pathway 110
Lung adenocarcinoma PP1 inhibits STAT3 signaling 111
PP5 Bladder cancer PP5 activates PPARγ signaling 108
PDP1 Various tumor cells PDP1 regulates glycolysis and TCA cycle 17, 83, 114
PDP2 Breast cancer PDP2 dephosphorylates ACSL4 109
ECM remodeling
and angiogenesis
PP2A Leukemia PP2A regulates MMPs expression 115
Endothelial cell PP2A regulates YAP activation 118
Calcineurin Endothelial cell Calcineurin/NFAT/angiopoietin-2 pathway promotes angiogenesis 120
PP4 CRC PP4 is associated with MMPs expression 19
PP5 Melanoma PP5 inhibits Hsp90α secretion 116
PPM1A Fibroblasts PPM1A regulates P38/MAPK pathway 82
Stromal cell interactions PP1 HCC PP1 regulates P38/MAPK pathway 20
AML PP1 regulates osteoblast-mediated protective effects 122
Various tumor cells PP1-MYPT1 activates YAP signaling 123, 124
Breast cancer, endothelial cell EV-derived PPP1R1B activates endothelial cells 125
PP2A HCC PPP2R2A regulates forkhead box O3, P27 and P21 8
Immune response PP1 Multiple myeloma PP1-GADD34 dissociation promotes immunogenic cell death 131
Various immune cells PP1-GADD34 mediates integrated stress response 23
B cell PP1 regulates to Fc clustering 156
Macrophage PP1 inhibits IRF3 signaling 157
DC PP1-GADD34 regulates IFN production 162
PP2A Various tumor cells PP2A inhibits STING signaling 18, 127
Pancreatic cancer, NSCLC Inflammatory stimuli regulate PP2A expression in tumor cells 133, 134
CD4+/CD8+ T cell PP2A suppresses T cell proliferation, glycolysis and cytokines production 6, 142, 144
Treg cell PP2A mediates maturation of Treg cells 149
B cell PP2A ensures optimal functions 153
TAM SET regulates metabolism, migration, M1 to M2 polarization and STING signaling 106, 158-160
MDSC PP2A terminates AKT/β-catenin pathway 161
Calcineurin Treg cell Calcineurin/NFAT pathway regulates Treg differentiation 70, 71
PP4 DC Calcineurin/NFAT pathway regulates cytokines production 163
OC PP4 regulates inflammatory factors secretion 128
T cell, lymphoma PP4-PP4R1 inhibits NF-κB signaling 147
Normal/leukemic T cell PP4 dephosphorylates PEA-15 148
PPM1B NSCLC PPM1B regulates NF-κB signaling 129
PPM1D HCC PPM1D is correlated with ICB inhibition 139
PPM1H CRC PPM1G is correlated with immune cell infiltration 138
PPM1K Pancreatic adenocarcinoma PPM1K is correlated with immune cell infiltration 81
breast invasive carcinoma PDP1 is negatively correlated with CD8+ T cells infiltration 140
PDP1 Breast invasive carcinoma PDP1 is negatively correlated with CD8+ T cells infiltration 140