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. 2024 Dec 24;16(1):46–71. doi: 10.1080/21645698.2024.2423987

Table 1.

Some orphan crops, their potential trait for crop improvement, research gaps and genome sequence status.

Crop [Common name) Scientific name Potential Genomic Markers Genome Sequence Status Chromosome Number Genome Size Breeding Effort Available Genomic Tools Availability
Transformation system
Research Gap References
Hyacinth bean Lablab purpureus L. Drought and salinity tolerance, nutritious, nutraceutical properties EST-SSRs, RAPD, EST-SNP, InDel. No 2 n = 2x = 24 367Mb Improved photoperiod insensitive and development of determinate pureline varieties Linkage map, QTL mapping Nil No superior cultivars and resistant varieties to pest and weeds 44,45
Zombi pea Vigna vexillata [L.] A. Rich Nutritious, biotic and abiotic stresses resistance RAD-seq, SSR, SNP No 2 n = 2x = 22 803Mb Identification of candidate gene, VvTaXI for Bruchid resistance QTL mapping Nil No improved zombi pea for abiotic stress resistance 24,46,47
African Yam Bean [Sphenostylis stenocarpa [Hochst ex. A. Rich.] Harms] Nutritious, versatile AFLP, ISSR, SSR, SNP 77–81% 2 n = 2x = 22 > 649Mb Identification of various agronomic traits Association mapping Nil There is need to improve crop yield, shorten the cooking and maturity periods, and increase the crop’s resilience to pests and diseases. 48,39
Winged bean Psophocarpus tetragonolobus (L.) DC. Nutritious, versatile, adaptable to diverse soil types AFLP, RAPD, SSR, SNP, 98.9% 2 n = 2x = 18 1.22Gb Identification of quantitative trait loci [QTLs) for traits such as pod and seed characteristics Cloning, transcriptomics, Genetic linkage maps Nil Little or no information on how to improve the long maturation period, indeterminate growth habit, low seed yield, and the need for a stake to support the vigorously growing vines 19,49
Bambara groundnut Vigna subterranea [L.] Verdc Nutritious, climate-resilient, versatile, adaptable to diverse environment ISSR, SSR, SNP No 2 n 2x = 22 550Mb Identification of QTL for some agronomic traits Whole Genome Sequencing, Transcriptome sequencing Nil No improved line for yield and adaptation to moisture stress 44,50,51
Finger millet Eleusine coracana Resilience to adverse climatic conditions, high nutritional content, versatile EST-SSR, ISSR, SSR, SNP 100% 2 n = 4x = 36 1.45Gb Development of lines with improved agronomic traits Linkage mapping, QTL mapping, Whole Genome sequencing, Reference genome, Transcriptomic, Proteomics, RNA-Seq Agrobacterium-mediated transformation and particle bombardment [biolistics] Identification of novel genes for abiotic stress tolerance and nutritional traits 52,53
Chickpea Cicer arietinum Nutritious, adaptability to diverse climates AFLP, ISSR, InDel, ISM, ILP, SSR, SNP 73.8% 2 n = 2x = 16 738Mb Identified loci associated with yield, drought tolerance, and resistance to diseases Linkage mapping, QTL mapping, GWAS, Whole Genome sequencing, Reference genome, Proteomics, RNA-Seq, DGE, CRISPR/Cas9 Agrobacterium-mediated transformation and particle bombardment [biolistics] Selection of accessions that can be used as parental material in breeding programs.
Lack of drought-tolerant cultivars.
54,55
Foxtail millet Setaria italica High nutritional content, environmental resilience, and potential health benefits KASP, AFLP, RAPD, InDel, SSR, SNP 100% 2 n = 2x = 18 490Mb Identification of genetic loci associated with drought resistance, plant height, flowering time, grain yield, and nutritional quality Linkage mapping, QTL mapping, GWAS, Whole genome Sequencing, Comparative Genomics, CRISPR Cas9, DGE, TILLING, MAS, GS Agrobacterium-mediated transformation and particle bombardment [biolistics] Lack of high yielding varieties 56,57
Grass pea Lathyrus sativus Ability to thrive in adverse conditions, high nutritional value AFLP, RAPD, EST-SSR, KASP, SSR, SNP 100% 2 n = 2x = 14 6.3Gbp Identification of stress-responsive genes Draft Genome Assemblies, Transcriptome sequencing [RNA-Seq], Comparative Genomics, Gene Knockout, TILLING Nil Lack of genetic improvement through GWAS and MAS 58,59
Horsegram Macrotyloma uniforum Nutritional and therapeutic properties, resilience to harsh environmental conditions RAPD, ISSR, SSR, SNP, ILP, COS 83.53% 2 n = 2x = 20 or 22 400Mb Identification of genes involved in metabolic process, Identification of candidate gene involved in environmental adaptation Linkage mapping, QTL mapping, Whole Genome Sequence, Transcriptome sequencing, Comparative Genomics, GWAS, DEG Nil Candidate genes underlying functional traits has not been identified 60,61
Pigeon pea Cajanus cajan Nutritious, versatile, nutraceutical properties, adaptability to marginal environments, drought resistance SSAP, REMAP, SCAO, SSR, SNP 100% 2 n 2x = 22 833Mb Identification of regions associated with Fusarium wilt resistance, sterility mosaic disease resistance, drought tolerance, and pod yield Whole Genome sequence, GWAS, MAS, Transcriptomics, Comparative Genomics, CRISPR Agrobacterium-mediated transformation Lack of breeding for high yielding pure line and hybrid 62,63
Moth bean Vigna aconitifolia Nutritious, adaptability to harsh climates, nutraceutical properties AFLP, RAPD, ISSR, SSR, SNP 50% 2 n 2x = 22 409Mb Identification of stress-responsive genes Linkage mapping, Proteomics, Comparative Genomics, Nil Lack of reference genome, little integration of modern genomics tools, Lack of common features of nutritional, anti-nutritional compounds, processing characteristics, and health benefits 64,65
Black gram Vigna mungo High protein content, adaptability to various cropping systems AFLP, RAPD, ISSR
, SSR, SNP
79.2% 2 n = 2x = 22 574Mb Identification of SNPs associated with seed size, flowering time, and biotic stress resistance Draft Genome Assembly, Linkage mapping, GWAS, QTL mapping, DEGs Agrobacterium-mediated transformation Incomplete reference genome, lack of high yielding and MYMV resistant varieties 66,67
Adzuki bean Vigna angularis Nutritious, functional food and medicinal applications AFLP, ISSR, EST-SSR, SSR, SNP 97.8% 2 n = 2x = 22 542Mb Identification of SNPs associated with seed size, flowering time, and disease resistance Draft Genome Assembly, Linkage mapping, GWAS, QTL mapping, DEGs, Comparative Genomics, MAS Agrobacterium-mediated transformation Little knowledge on utilization.
Little understanding of the anti-obesity mechanism and the specific bioactive components involved has not been identified.
68–70
Mung bean Vigna radiata Nutritious, nutraceutical properties, drought tolerance, disease resistance, climate-resilient AFLP, RAPD, ISSR, SSR, SNP 96.9% 2 n = 2x = 22 543Mb Identification of SNPs associated with seed size, flowering time, and disease resistance Draft Genome Assembly, Linkage mapping, GWAS, QTL mapping, DEGs, Comparative Genomics, MAS Agrobacterium-mediated transformation Little knowledge on the unraveling of the main functional components relevant to health benefits 71,72
Fonio Millet Digitaria exilis Fast maturing, climate-resilient, nutritious, nutraceutical properties, highly adaptable to poor soils and drought conditions AFLP, RAPD, SSR, SNP 99% 2 n = 4x = 36 761Mb Identification of drought resistance traits GWAS, DEGs, Comparative Genomics Nil Improvement and domestication is lacking 73