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. 2025 Jan 17;13:RP95273. doi: 10.7554/eLife.95273

Table 1. Definition of handcrafted features (according to the scikit-image documentation).

All of these features were extracted for each fluorescent channel (1-4) and region (nucleus, cytoplasm, and whole-cell) in the cell painting images.

Feature name Description Category
Intensity max Maximal pixel intensity inside the ROI Intensity
Intensity mean Mean pixel intensity inside the ROI Intensity
Intensity min Minimal pixel intensity inside the ROI Intensity
Intensity Std Standard deviation of the pixel intensities inside the ROI Intensity
Area Area of the ROI Shape
Area convex The area of the convex polygon that encloses the ROI Shape
Area filled Area of the ROI including all holes inside the ROI Shape
Axis major length The length of the major axis of the ellipse that has the same normalized second central moments as the region. Shape
Axis minor length The length of the minor axis of the ellipse that has the same normalized second central moments as the region. Shape
Centroid Coordinates of the center of the ROI Shape
Eccentricity Eccentricity of the ellipse that has the same second-moments as the ROI. Shape
Equivalent diameter area The diameter of a circle with the same area as the ROI. Shape
Extent The ratio of the Area of the ROI to the Area of the bounding box around the ROI. Shape
Feret diameter max The longest distance between points around a ROIs’ convex polygon. Shape
Orientation The angle between the X-axis and the major axis. Shape
Perimeter Length of the contour of the ROI Shape
Perimeter crofton Perimeter of the ROI approximated by the Crofton formula in four directions. Shape
Solidity The ratio of the number of pixels inside the ROI to the number of pixels of the convex polygon. Shape
Contrast The difference between the maximum intensity and minimum intensity pixel inside the ROI. Texture
Dissimilarity The average difference in pixel intensity between neighbouring pixels. Texture
Homogeneity Value for similarity of pixels in the ROI. Texture
Energy Value for the local change of pixel intensities in the ROI. Texture
Correlation Value for the linear dependency of pixels in the ROI. Texture
ASM (angular second moment) Value for the uniformity of pixel values in a ROI. Texture