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. 2025 Apr 14;16:3525. doi: 10.1038/s41467-025-58506-x

Table 4.

Summary of top variants in the APOC1 gene cluster exhibiting genome-wide significance for the TyG index, a marker of insulin resistance, identified within the Taiwan Biobank cohort

GENE CHR Interval (b38) Top SNP Pos (b38) Top P Novel TWAS GWAS catalog
NECTIN2 19 44846297..44889223 rs283811 44,885,243 1.6E-94 yes 4 TG, HDL-C, T2D, AD, BMI
TOMM40 19 44891254..44903689 rs157582 44,892,962 7.5E-94 yes NA TG, HDL-C, AD, BMI
APOE 19 44905796..44909393 rs440446 44,905,910 1.7E-106 yes 1 TG, HDL-C, T2D, AD, BMI
APOC1 19 44914325..44919346 rs5117 44,915,533 2.8E-111 no 8 TG, HDL-C, AD, BMI, TG:HDL-C
APOC1P1 19 44926803..44931386 rs7259004 44,929,300 2.0E−59 no NA TG, HDL-C, AD, BMI, TG:HDL-C
APOC2, APOC4-APOC2 19 44946051..44949565 rs2288911 44,946,027 2.8E-10 yes 3, 10 TG, HDL-C, AD
TAD CHR Interval (b38) -- -- -- -- -- --
pancreas 19 43760000..44560000 -- -- -- -- -- --
pancreas 19 45440000..47760000 -- -- -- -- -- --
hippocampus 19 43520000..44720000 -- -- -- -- -- --
hippocampus 19 45640000..47720000 -- -- -- -- -- --
liver 19 42080000..44560000 -- -- -- -- -- --
liver 19 45000000..47720000 -- -- -- -- -- --

Tissues in TWAS: 1 = Adipose Subcutaneous; 2 = Adipose Visceral Omentum; 3 = Adrenal Gland; 4 = Brain Cortex; 5 = Brain Hippocampus; 6 = Brain Hypothalamus; 7 = Liver; 8 = Muscle Skeletal; 9 = Pancreas; 10 = Pituitary; 11 = Thyroid. Previous GWASs reported in the NHGRI-EBI GWAS Catalog have identified various phenotypes that reach genome-wide significance (P < 5 × 10−8). Coordinates for TADs were retrieved from the following source (http://3dgenome.fsm.northwestern.edu/publications.html).

AD Alzheimer’s disease, b38 Genome Research Consortium human build 38, BMI body mass index, CHR chromosome, FG fasting glucose, HDL-C high-density lipoprotein cholesterol, IR insulin resistance, NA not available, Pos position, T2D type 2 diabetes, TAD topologically associating domains, TG triglycerides, TG:HDLC the triglyceride to high-density lipoprotein cholesterol ratio, TWAS transcriptome-wide association analysis.