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. 2025 May 8;15:16061. doi: 10.1038/s41598-025-99485-9

Table 2.

Spearman correlations of the Microbiome and parameter changes between timepoints 6 h and 24 h after Bruce intervention. Only correlations with significant p-values (< 0.05) are included.

Group Parameter Microbiome feature Spearman correlation Adjusted p value
Control Leptin [pg/ml] PWY0-1241: ADP-l-glycero-&beta;-d-manno-heptose biosynthesis 0.95 < 0.001
LIF [pg/ml] RIBOSYN2-PWY: flavin biosynthesis I (bacteria and plants) 0.90 0.02
IL-6 [pg/ml] Bacteroides cellulosilyticus − 0.88 0.03
LIF [pg/ml] PWY-5971: palmitate biosynthesis (type II fatty acid synthase) − 0.87 0.06
Il-10 [pg/ml] PWY0-1241: ADP-l-glycero-&beta;-d-manno-heptose biosynthesis − 0.83 0.10
LIF [pg/ml] Bacteroides caccae − 0.84 0.11
LIF [pg/ml] Clostridiaceae bacterium 0.80 0.12
LIF [pg/ml] BIOTIN-BIOSYNTHESIS-PWY: biotin biosynthesis I − 0.80 0.12
LIF [pg/ml] FASYN-ELONG-PWY: fatty acid elongation—saturated − 0.79 0.12
LIF [pg/ml] PWY-7664: oleate biosynthesis IV (anaerobic) − 0.79 0.12
LIF [pg/ml] PWY-6609: adenine and adenosine salvage III 0.83 0.12
LIF [pg/ml] PWY-6519: 8-amino-7-oxononanoate biosynthesis I − 0.79 0.12
LIF [pg/ml] PWY0-862: (5Z)-dodecenoate biosynthesis I − 0.79 0.12
LIF [pg/ml] PWY-6282: palmitoleate biosynthesis I (from (5Z)-dodec-5-enoate) − 0.79 0.12
Leptin [pg/ml] Roseburia faecis − 0.82 0.15
LIF [pg/ml] PWY-6572: chondroitin sulfate degradation I (bacterial) − 0.78 0.15
FSTL1 [pg/ml] PWY0-1241: ADP-l-glycero-&beta;-d-manno-heptose biosynthesis − 0.80 0.16
Il-10 [pg/ml] PWY-6700: queuosine biosynthesis I (de novo) 0.80 0.17
LIF [pg/ml] PWY-8187: L-arginine degradation XIII (reductive Stickland reaction) − 0.76 0.18
SPARC [pg/ml] PWY0-1241: ADP-l-glycero-&beta;-d-manno-heptose biosynthesis − 0.80 0.18
Strength IL-15 [pg/ml] PWY0-1298: superpathway of pyrimidine deoxyribonucleosides degradation − 0.95 < 0.02
IL-10 [pg/ml] Clostridium sp AF27 2AA 0.92 0.10
IL-6 [pg/ml] Gemmiger formicilis 0.92 0.11
Endurance IL-15 [pg/ml] Eubacterium rectale − 0.95 < 0.001
TfR [pg/ml] Blautia wexlerae − 0.93 < 0.01
TIMP1 [pg/ml] Bacteroides thetaiotaomicron 0.92 < 0.01
TIMP1 [pg/ml] PWY-5188: tetrapyrrole biosynthesis I (from glutamate) − 0.92 < 0.01
Adiponectin [pg/ml] PWY3O-4107: NAD salvage pathway V (PNC V cycle) − 0.92 < 0.01
Adiponectin [pg/ml] PWY-4041: &gamma;-glutamyl cycle − 0.90 < 0.01
TIMP1 [pg/ml] Blautia wexlerae − 0.90 < 0.01
Adiponectin [pg/ml] COLANSYN-PWY: colanic acid building blocks biosynthesis − 0.89 < 0.01
TIMP1 [pg/ml] GLYCOGENSYNTH-PWY: glycogen biosynthesis I (from ADP-D-Glucose) − 0.88 < 0.01
TIMP1 [pg/ml] PWY-5030: l-histidine degradation III 0.88 < 0.01
TIMP1 [pg/ml] HISDEG-PWY: l-histidine degradation I 0.88 < 0.01
TIMP1 [pg/ml] Odoribacter splanchnicus − 0.88 < 0.01
TIMP1 [pg/ml] Bacteroides uniformis 0.84 < 0.01
TIMP1 [pg/ml] Dorea longicatena 0.84 < 0.01
TIMP1 [pg/ml] Lacrimispora celerecrescens − 0.84 < 0.01
TIMP1 [pg/ml] OANTIGEN-PWY: O-antigen building blocks biosynthesis (E. coli) − 0.84 < 0.01
TIMP1 [pg/ml] DAPLYSINESYN-PWY: l-lysine biosynthesis I − 0.83 < 0.01
TIMP1 [pg/ml] CITRULBIO-PWY: L-citrulline biosynthesis 0.83 < 0.01
TfR [pg/ml] KETOGLUCONMET-PWY: ketogluconate metabolism − 0.82 0.12
TfR [pg/ml] PWY-5188: tetrapyrrole biosynthesis I (from glutamate) − 0.88 0.12
TIMP1 [pg/ml] PWY-7199: pyrimidine deoxyribonucleosides salvage 0.78 0.16
TIMP1 [pg/ml] NONOXIPENT-PWY: pentose phosphate pathway (nonoxidative branch) I − 0.78 0.16
TIMP1 [pg/ml] Eubacterium rectale − 0.78 0.16
TIMP1 [pg/ml] HSERMETANA-PWY: l-methionine biosynthesis III − 0.78 0.16
TIMP1 [pg/ml] P441-PWY: superpathway of N-acetylneuraminate degradation − 0.84 0.17
TIMP1 [pg/ml] ARGSYNBSUB-PWY: l-arginine biosynthesis II (acetyl cycle) − 0.77 0.17
TIMP1 [pg/ml] METH-ACETATE-PWY: methanogenesis from acetate − 0.77 0.17
TIMP1 [pg/ml] ARGSYN-PWY: l-arginine biosynthesis I (via l-ornithine) − 0.77 0.17
TfR [pg/ml] PWY-5030: l-histidine degradation III 0.84 0.18
TfR [pg/ml] Eubacterium rectale − 0.84 0.18
TIMP1 [pg/ml] PWY-7237: myo-, chiro- and scyllo-inositol degradation − 0.76 0.19
TIMP1 [pg/ml] P41-PWY: pyruvate fermentation to acetate and (S)-lactate I − 0.76 0.19
TIMP1 [pg/ml] PWY-6823: molybdopterin biosynthesis − 0.76 0.19

TIMP1 (tissue inhibitor of metalloproteinases), IL-5 (interleukin 5), IL-6 (interleukin 6), IL-10 (interleukin 10), IL-15 (interleukin15), LIF (leukemia inhibitory factor), SPARC (secreted protein acidic and cysteine rich), TfR (transferrin receptor). Significant correlations (p-value < 0.05) were observed across all groups. In the control group, ADP-L-glycero-β-D-manno-heptose biosynthesis was positively correlated with leptin, while Bacteroides cellulosilyticus showed negative correlations with IL-6. The strength group revealed strong associations betweenof pyrimidine deoxyribonucleosides degradation and IL-15. In the endurance group, notable correlations included Eubacterium rectale with IL-15, and Blautia wexlerae with TfR, as well as various pathways involved in NAD (Nicotinamide Adenine Dinucleotide) salvage and histidine degradation. These findings highlight the dynamic microbiome-blood marker interactions induced by the Bruce test across different exercise modalities.

Significant values are in bold.