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. 1998 Apr;18(4):2067–2076. doi: 10.1128/mcb.18.4.2067

FIG. 8.

FIG. 8

A possible pathway for multistep evolution of a new, tissue-specific gene promoter. Layering of positive (+) and negative (−) elements with some tissue preference can result in high levels of tissue-specific expression of a recruited gene in the lens.