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. 2025 Aug 21;8:537. doi: 10.1038/s41746-025-01911-9

Table 3.

List of applied clinical and omics ageing clocks

Clock Reference Data type Remarks
MLR-Levine28 Levine, ME. 2013. Gerontol A Biol Sci Med Sci. Clinical Predicts chronological/ biological age
PCA-Levine28 Predicts chronological/ biological age
KDM31 Klemera, P. and Doubal, S. 2006. Mech Ageing Dev.
ProtAge-Tanaka30 Tanaka, T., et al. 2018. Aging Cell. Proteomics Predicts chronological/ biological age
ProtAge-MD29 Macdonald-Dunlop, E., et al. 2022. Aging. Predicts chronological/ biological age
ClinAge-MD29 Macdonald-Dunlop, E., et al. 2022. Aging. Clinical Overestimated BA (median BA 750 years), not shown
MetaboAge-MD29 Macdonald-Dunlop, E., et al. 2022. Aging. Metabolomics Overestimated BA (median BA 600 years), not shown
MetaboAge-vdA32 van den Akker, EB., et al. 2020. Circ Genom Precis Med. Underestimated BA (median BA –5000 years), not shown
Metabonomics33 Hertel, J., et al. 2016. Journal of Proteome Research. Missing variables (only 22% vars. are available), not shown
Skin & blood clock52 Horvath, S., et al. 2018. Aging. DNA methylation Predicts chronological age
MethylDetectRAge43 Hillary, R. and Marioni, R. Wellcome Open Res. Predicts chronological age
Multi-tissue clock53 Horvath, S. 2013. Genome Bioloy. Predicts chronological/biological age
Hannum clock26 Hannum, G., et al. 2013. Molecular Cell. Predicts chronological/biological age
mPoA25 Belsky, D., et al. 2020. eLife. Predicts the pace of ageing
DunedinPACE56 Belsky, D., et al. 2022. eLife. Predicts the pace of ageing
PhenoAge54 Levine, M., et al. 2018. Aging. Predicts health- and lifespan
GrimAge27 Lu, A., et al. 2019. Aging. Predicts health- and lifespan
GrimAge255 Lu, A., et al. 2022. Aging. Predicts health- and lifespan
Metaclock34 Liu, Z., et al. 2020. Aging Cell. Predicts mortality
EpiTOC35 Yang, Z., et al. 2016. Genome Biology. Predicts mitotic age
EpiTOC236 Teschendorff, A.E. 2020. Genome Medicine Predicts mitotic age
MiAge37 Youn, A., and Wang, S. 2018. Epigenetics. Predicts mitotic age
DNAmTL38 Lu, A., et al. 2019. Aging. DNA methylation Predicts telomere length
CausAge61 Ying, K, et al. 2024. Nature Aging. Predicts chronological/ biological age
DamAge61 Ying, K, et al. 2024. Nature Aging. Predicts chronological/ biological age
AdaptAge61 Ying, K, et al. 2024. Nature Aging. Predicts chronological/ biological age
GOLD BioAge71 Hao, et al. 2025. Adv. Sci. Clinical Predicts morbidity & mortality
GOLD ProtAge71 Metabolomics Predicts morbidity & mortality
GOLD MetAge71 Proteomics Predicts morbidity & mortality

Rows with double dagger symbol (‡) refer to clocks that are further investigated.