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. 2025 Aug 22;87(9):134. doi: 10.1007/s11538-025-01509-y

Table 2.

Definitions of binary imbalance indices, which are only applicable to binary trees, i.e., those with domain BTn∗. It is straightforward to see that these binary imbalance indices are induced by the balance value metaconcept ΦfB when the function f is chosen as specified in the right column. The (quadratic) Colless index is induced by the first-order metaconcept where id is the identity function, while the corrected Colless index is induced by the second-order metaconcept.

binary imbalance index definition BVM ΦfB
Colless index Colless (1982); Shao and Sokal (1990) CT:=∑v∈V˚TbT(v) id
Corrected Colless index Heard (1992) ICT:=2(n-1)(n-2)·CT; ICT:=0 for n=1,2 fIC(b,n)=2(n-1)(n-2)·b
Quadratic Colless index Bartoszek et al. (2021) QCT:=∑v∈V˚TbT(v)2 fQC(b)=b2