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. 2025 Aug 22;87(9):134. doi: 10.1007/s11538-025-01509-y

Table 2.

Definitions of binary imbalance indices, which are only applicable to binary trees, i.e., those with domain BTn. It is straightforward to see that these binary imbalance indices are induced by the balance value metaconcept ΦfB when the function f is chosen as specified in the right column. The (quadratic) Colless index is induced by the first-order metaconcept where id is the identity function, while the corrected Colless index is induced by the second-order metaconcept.

binary imbalance index definition BVM ΦfB
Colless index Colless (1982); Shao and Sokal (1990) CT:=vV˚TbT(v) id
Corrected Colless index Heard (1992) ICT:=2(n-1)(n-2)·CT; ICT:=0 for n=1,2 fIC(b,n)=2(n-1)(n-2)·b
Quadratic Colless index Bartoszek et al. (2021) QCT:=vV˚TbT(v)2 fQC(b)=b2