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. 2025 Aug 22;12:1467. doi: 10.1038/s41597-025-05793-0

Table 1.

Environmental variables selected for modeling.

Variables Description Period & Time Interval Resolution Source
Bio1 Annual mean temperature

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio2 Mean diurnal temperature range

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio3

Isothermality

(Bio2/Bio7)(×100)

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio4

Temperature seasonality

(standard deviation × 100)

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio5 Maximum temperature of warmest month

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio6 Minimum temperature of coldest month

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio7 Temperature annual range

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio8 Mean temperature of wettest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio9 Mean temperature of driest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio10 Mean temperature of warmest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio11 Mean temperature of coldest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio12 Annual precipitation

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio13 Precipitation of wettest month

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio14 Precipitation of driest month

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio15

Precipitation seasonality

(Coefficient of Variation)

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio16 Precipitation of wettest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio17 Precipitation of driest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio18 Precipitation of warmest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Bio19 Precipitation of coldest quarter

Historical, 30 years mean (1970–2000);

Future, 20 years mean (2021–2100)

1/12° WorldClim
Elevation

Altitude above sea level

(Topographic Position)

Constant 1/12° WorldClim
Slope

Rate of elevation change across space

(Terrain Steepness)

Constant 1/12° Calculated
NDVI Normalized Difference Vegetation Index Historical, 18 years mean (1982–2000) 1/12° PKU