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. 2025 Aug 25;22:23. doi: 10.1186/s12983-025-00569-z

Table 2.

Effects of female traits and climatic variables on the number of fledglings

Sources of variation Estimate SE z P
Conditional model (N = 7315 broods of 5513 females)
Intercept 1.52 0.02 69.11  < 0.001
Female age − 0.01 0.01 − 0.83 0.41
Female body condition 0.00 0.01 − 0.13 0.90
Within-year temperature: incubation 0.00 0.01 0.43 0.67
Between-year temperature: incubation 0.01 0.02 0.47 0.64
Within-year precipitation: incubation 0.00 0.01 0.16 0.87
Between-year precipitation: incubation 0.03 0.02 1.71 0.088
Within-year temperature: nestlings 0.02 0.01 1.44 0.15
Between-year temperature: nestlings − 0.02 0.02 − 0.86 0.39
Within-year precipitation: nestlings − 0.01 0.01 − 1.79 0.073
Between-year precipitation: nestlings − 0.04 0.02 − 1.69 0.092
Laying date − 0.07 0.02 − 4.07  < 0.001
Clutch size 0.09 0.01 12.38  < 0.001
Female ID random 0.00
Plot ID random 0.05
Year of study random 0.10
Pseudo-R2marginal/conditional 0.08/0.13
Zero-inflated model (N = 7315 broods of 5513 females)
Intercept − 1.96 0.18 − 11.03  < 0.001
Female age 0.07 0.08 0.87 0.38
Female body condition 0.04 0.04 1.16 0.24
Within-year temperature: incubation − 0.07 0.05 − 1.47 0.14
Between-year temperature: incubation − 0.28 0.19 − 1.44 0.15
Within-year precipitation: incubation − 0.02 0.04 − 0.53 0.60
Between-year precipitation: incubation − 0.19 0.14 − 1.32 0.19
Within-year temperature: nestlings − 0.15 0.04 − 3.45  < 0.001
Between-year temperature: nestlings 0.25 0.20 1.22 0.22
Within-year precipitation: nestlings 0.12 0.04 2.94 0.003
Between-year precipitation: nestlings 0.31 0.18 1.78 0.076
Laying date 0.73 0.08 8.94  < 0.001
Clutch size 0.11 0.04 2.86 0.004
Female ID random 0.00
Plot ID random 0.47
Year of study random 0.87

Output of the zero-inflated generalized linear mixed model with Poisson error distribution and the log-link function for a conditional component and with binomial error distribution and the logit-link function for a zero-inflated component testing how female age (categorical predictor), female body condition, the within- and between-year effects of ambient temperature and sum of precipitation experienced during the incubation and nestling period, laying date, and clutch size (all as continuous predictors), affect the number of fledglings. All continuous explanatory terms were standardized. The female identity, study plot identity, and year of study were included as random factors. Significant terms P < 0.05 are in bold