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. 2025 Aug 12;12:1562641. doi: 10.3389/fvets.2025.1562641

Table 12.

Antioxidant effects of THY across multiple species and models.

Species/ model THY Dosage/ formulation Key measurements Findings References Year
Aquatic species
Nile tilapia 1–2 mL/kg feed MDA, GSH reductase in muscle/liver, catalase Reduces MDA, enhances antioxidant enzyme activity (2) 2018
Thyme powder (THP; 2%) with insecticide exposure MDA in O. niloticus Reduces MDA levels compared to control; mitigates insecticide-induced oxidative damage (13) 2020
Rainbow trout Phytogenic additive (6 g/kg of THY) MDA, GSH-based enzymes, catalase Reduces MDA, increases GSH activity, increases catalase (114) 2012
0, 500, 1,000, 1,500 mg/kg THEO SOD, GPx, MDA Increases liver SOD and GPx, reduces MDA (115) 2015
THEO ALT, AST, cortisol, glucose, CAT, GPx THY suppresses oxytetracycline-induced enzyme changes; increases gut CAT and GPx activities (116) 2018
Common carp THY (short/long exposure) MDA, oxidative stress markers Lower MDA, less stress-induced oxidative markers with THY (117) 2018
Mammalian models
Mice THY (60 mg/kg in DSS model) MPO, MDA, GSH, SOD Loweres MPO and MDA, increases GSH and SOD in colonic tissues, indicating antioxidant effect in DSS colitis model (45) 2018
THY (0, 20, 40, 80 mg/kg) Lung MDA THY dose-dependently reduces LPS-induced MDA in lung tissue (108) 2018
Rats THEO Lipid peroxidation, SOD, GSH Reduces lipid peroxidation and restores SOD and GSH levels in T. vitulorum-infected rats (118) 2013
THEO (5.6% THY) and olive EOs (1.5 g/kg bw) SOD, GPx, CAT, non-enzymatic antioxidant capacity Increases catalase, reduces SOD and GPx, modulates plasma antioxidant capacity; enhances bioavailability of olive phenolics in combination (119) 2014
THY oral administration TAC, TOC, TBARS, GSH Reverses hydrocortisone-induced oxidative stress markers, elevates TAC, and decreases TOC (106) 2015
Rabbits THY (3–6 mg/kg/day) MDA, serum TAC Lowers MDA and enhances TAC in the serum of high-fat diet animals (112) 2016
THY (250 mg/kg feed) MDA, GPx, LDH Reduces MDA and GPx in blood; decreases LDH in muscle tissue (120) 2020
Pigs EOs blend (18% THY, 0.01%) Plasma TAC Increases TAC (18) 2012
EOs blend (13.5% THY, 4.5% CIN) Plasma antioxidant capacity Increases plasma antioxidant capacity (121) 2014
THY-CAR blend (1:1, 100 mg/kg) SOD, GPx, TBARS in jejunum Elevates SOD, GPx, and reduces TBARS in jejunum in supplemented group (15) 2017
THY (0.01% in diet) SDH, LDH, drip loss Reduces oxidative markers, enhances SDH activity, and improves muscle quality (122) 2020
EOs blend (10% THY, 10% CIN) Hepatic SOD, GSH Higher hepatic SOD activity; no reduction in GSH, suggesting stimulated antioxidant defense (123) 2023
Dairy cows THEO (THY ~59%) NF-κB binding energy Demonstrates potential against oxidative stress, with binding energy of THY to NF-κB indicating suitability for oxidative stress management (124) 2023
Poultry
Broiler chickens THY-CAR blend (0, 60, 100, and 200 mg/kg) SOD, GPx, MDA Increases SOD, GPx; decreases MDA in liver, serum, and thigh muscle (125) 2013
THY (200 mg/kg) Plasma MDA Significantly reduces MDA in plasma (22) 2014
Nigella seed EO (0, 0.5, 1, and 1.5 g/kg) SOD, GSH, GPx Greater SOD and GSH with 1.5 g/kg; increases GPx in antibiotic and 1 g/kg groups (21) 2015
THY (250 mg/kg feed) Serum GPx Elevates GPx in heat-stressed chickens (126) 2016
THEO (0, 0.05, and 0.1%) SOD, MDA Elevates SOD and reduces MDA levels in plasma with THEO supplementation (59) 2019
EOs blend (0–400 mg/kg) SOD, MDA, GPx, total antioxidant capacity SOD increases in higher doses (200–400 mg/kg); no significant effect on MDA, GPx, and total antioxidant capacity (19) 2021
Turkey THY (0, 30 mg/kg) or EO blend Lipid oxidative stability, GSH enzymes Improves lipid oxidative stability and increases GSH-based enzyme activity (127) 2014
Laying hens THP (0–9 g/kg) SOD, GSH, MDA Increases SOD, GSH; reduces MDA (128) 2015
THP (0.9%) SOD, MDA Increases SOD, decreases MDA levels (1) 2017
In vitro and ex vivo studies
Porcine brain tissue THEO from three different Thymus species (THY: 38.5%, 49.10%, and 56.02%) Radical scavenging, thiobarbituric acid assay Highest radical scavenging in T. serpyllum, followed by T. algeriensis, and T. vulgaris Eos (32) 2013
RAW264.7 macrophages THY (10, 20, and 40 μg/mL) ROS Dose-dependent inhibition of LPS-induced ROS production (41) 2017
IPC-J2 cells THY pretreatment (0, 50 μM) ROS Inhibites LPS-induced ROS production (25) 2019
Bovine ovarian tissue THY (400, 800, 1,600, and 3,200 μg/mL) mRNA for SOD1, CAT, GPX1, PRDX6 Increases CAT activity at 800 μg/mL; reduces mRNA expression of SOD1, CAT, PRDX6 at 400–800 μg/mL (129) 2024