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. 2025 Sep 4;16:1638147. doi: 10.3389/fphar.2025.1638147

TABLE 5.

Antimicrobial categories of S. rebaudiana.

Category Microorganism (Latin) Extract type Active results Antimicrobial index (e.g., MIC/zone) Mechanism (if known) Ref.
bacteria Escherichia coli O157 Fermented hot water extract Exhibited bactericidal activity Complete killing at 40% (v/v) Acidic pH-dependent bactericidal effect; suppression of VT1/VT2 toxin production Ruiz-Ruiz et al. (2017)
Tomita et al. (1997)
Enterotoxigenic Escherichia coli EC127 Fermented hot water extract Viable count reduced from 109 to <105 cfu/mL within 2 h at 20% (v/v) High susceptibility Acidic pH disturbs extracellular environment and interferes with metabolic activity
Enteroinvasive Escherichia coli EC174 Fermented hot water extract Viable count reduced from ∼109 to <102 cfu/mL within 2 h at 20% (v/v) High susceptibility at 20%; total kill at 40% (v/v) Acidic pH damages the cell membrane and leads to cell death
Escherichia coli Crude methanolic extract Exhibited bactericidal activity Positive control: amoxicillin; negative: <1% DMSO; 69% inhibition rate -- (Tomita et al., 1997; Sumit et al., 2008; Maryam et al., 2015; Ruiz-Ruiz et al., 2017; Chakma et al., 2023; Stachurska et al., 2023)
Volatile extract (e.g., petroleum ether) Exhibited bactericidal activity; complete inhibition at low concentration MIC as low as 250 µg/mL --
Aqueous extract Inhibition observed 7 mm (low activity) --
Ethanol extract (shaking or Soxhlet method) Moderate to high antibacterial activity observed Inhibition zone range: 6.53–18.81 mm; max: 18.81 mm --
Escherichia coli DSM 5695 SRa (25%, acetone) Initial inhibition followed by rebound MIC: 25%; toxic Concentration-dependent toxicity; weakened under dynamic conditions Stachurska et al. (2023)
SRm (25%, Methanol) Enhanced lysis followed by rebound MIC: 25%; detoxified but activated Promotes lysis; growth stimulation under dynamic conditions
Escherichia coli DSM 613 SRa (25%, acetone) OD significantly reduced MIC: 25%; toxic Toxic under static, but neutralized under dynamic conditions
SRm (25%, acetone) No significant antibacterial activity None No apparent toxicity or inhibition
Proteus mirabilis Polar extracts (methanol, ethyl acetate, hexane) All extracts showed activity; hexane extract exhibited the strongest inhibition Moderate to high activity -- (Tomita et al., 1997; Sumit et al., 2008; Ruiz-Ruiz et al., 2017)
Chloroform Inhibition (10 mm zone of inhibition) 10 mm Antibacterial activity
Salmonella spp. Fermented hot water extract Viability reduced from ∼109 to <102 cfu/mL within 2 h at 20% (v/v) concentration High susceptibility Low pH disrupts bacterial metabolism and inhibits growth Tomita et al. (1997)
Staphylococcus aureus Crude methanolic extract Inhibitory rate of 50%; Amoxicillin used as positive control; DMSO (<1%) as negative control -- -- (Tomita et al., 1997; Sumit et al., 2008; Maryam et al., 2015; Ruiz-Ruiz et al., 2017; Chakma et al., 2023)
Shaking extraction (ethanol: water = 80:20), 50 mg/mL Highest inhibition zone reached 28.00 mm; range: 10.33–28.00 mm 10.33–28.00 mm; classified as high activity (>14 mm) --
Fermented hot water extract (30%–40% v/v) Significant inhibition after 2 h Approx. 30%–40% (v/v); moderately sensitive Acidic pH weakens cell wall synthesis, causing structural disruption
Pseudomonas aeruginosa SRa (25%, acetone extract of S. rebaudiana) No synergy with Phi6 phage; OD decreased under static condition MIC: 25%; growth inhibition under static conditions Weak toxicity; activity reduced under dynamic condition (Sumit et al., 2008; Ruiz-Ruiz et al., 2017; Chakma et al., 2023; Stachurska et al., 2023)
Shaking extraction (ethanol:water = 50:50), 25–50 mg/mL Inhibition zone up to 14.67 mm; range: 6.27–14.67 mm 6.27–14.67 mm; moderate activity (10–14 mm) --
Pseudomonas syringae SRm (25%, Methanol) No synergy with Phi6; flat growth curve -- Limited impact on growth; no lysis or synergy Stachurska et al. (2023)
Enterococcus faecalis Shaking extraction (ethanol:water = 50:50), 50 mg/mL Inhibition zone up to 18.00 mm; range: 6.67–18.00 mm 6.67–18.00 mm; moderate to high activity (≥14 mm) -- (Sumit et al., 2008; Chakma et al., 2023)
Bacillus subtilis Crude methanolic extract Inhibitory rate of 51.8%; Amoxicillin used as positive control; DMSO (<1%) as negative control Inhibitory rate: 51.8% (qualitative) -- (Tomita et al., 1997; Sumit et al., 2008; Maryam et al., 2015)
water extract Inhibition zone approximately 8–9 mm mild activity (8–9 mm) --
Vibrio parahaemolyticus Fermented hot water extract <10% (v/v) extract significantly reduced colony count within 2 h; highly sensitive Not numerically specified (highly sensitive) Extremely sensitive to acidic pH; low concentration causes cell lysis Tomita et al. (1997)
Bacillus cereus Fermented hot water extract <10% (v/v) extract showed significant inhibition, but spores may survive Not numerically specified; sensitive (spore resistance noted) Acidic pH affects vegetative cells; spores highly resistant but partially inactivated
Yersinia enterocolitica Fermented hot water extract 30%–40% (v/v) concentration caused significant inhibition within 2 h Not numerically specified (moderately sensitive) Acidic pH inhibits physiological activity and disrupts cell membrane integrity
Listeria monocytogenes Stevia ethanolic extract Inhibitory at 62.5–500 µg/mL; MIC = 125 µg/mL, MBC = 250 µg/mL; inhibited biofilm formation (P < 0.01), damaged membranes, caused nucleic acid leakage MIC = 125 µg/mL; MBC = 250 µg/mL Damages cell membrane, inhibits biofilm formation, induces nucleic acid leakage Belda-Galbis et al. (2014)
Fungi Candida albicans Water, ethanol, and acetone extracts Acetone extract showed strongest inhibition; more effective than water extract Inhibition zone: 17–21 mm (acetone); lower for other solvents Likely related to cell wall or membrane disruption (Jayaraman et al., 2008; Ruiz-Ruiz et al., 2017)
Trichophyton rubrum Leaf extract Maximum inhibition zone of 12 mm at a concentration of 7.5 µg/mL Inhibition zone: 9–12 mm Possibly disrupts cell membrane and/or cell wall integrity (Jayaraman et al., 2008; Sumit et al., 2008)
A.niger Petroleum Ether Inhibition (16 mm zone of inhibition 16 mm Antifungal activity Sumit et al. (2008)
Sumit et al. (2008)
P. chrysogenum Cyclohexane Inhibition (14 mm zone of inhibition) 14 mm Antifungal activity
A. solani Chloroform Inhibition (16 mm zone of inhibition) 16 mm Antifungal activity
Viruses Rotavirus Methanolic leaf extract Maximum inhibition ∼71.6% at 200 µg/mL; no significant cytotoxicity observed IC50 ≈ 95 µg/mL; CC50 > 300 µg/mL May interfere with viral adsorption/entry; activates host defenses Takahashi et al. (2001)
Herpes Simplex Virus Type 1 Polysaccharide fractions (SFW, SSFK) Both fractions significantly inhibited HSV-1 infection. SFW: EC50 = 0.3 µg/mL, SI = 917; SSFK: EC50 = 18.8 µg/mL, SI > 53. Reduced plaque size post-entry. SFW: EC50 = 0.3 µg/mL; SSFK: EC50 = 18.8 µg/mL; SI > 50 Interferes with viral adsorption, penetration, glycoprotein expression, and cell-to-cell transmission; heparin-mimetic competition (Ceole et al., 2020; Aghababaei and Hadidi, 2023)
Parasites Trypanosoma cruzi DCM extract of Stevia aristata 100 µg/mL killed 100% of parasites in vitro within 96 h; EC50 = 47.9 µg/mL; significantly reduced cyst weight in vivo (5.47 → 2.55 g, p < 0.05) EC50 = 47.9 µg/mL; total kill at 100 µg/mL (96 h) -- Albani et al. (2022)
Entamoeba histolytica Stevioside (STV) IC50 = 9.53 mM (24 h); induced cell membrane irregularity, autophagic vacuole formation, and morphological abnormalities; reduced liver damage in vivo IC50 = 9.53 mM (24 h) Disrupts membrane structure, induces autophagy-like vacuoles, inhibits cysteine protease expression and proteolytic activity Ortega-Carballo et al. (2024)

S. rebaudiana acetone extract → SRa.

S. rebaudiana methanol extract → SRm.

Activity level based on inhibition zone diameter.

  <7 mm: No activity.

  7–10 mm: Low activity.

  10–14 mm: Moderate activity.

  14 mm: High activity.

Antibacterial Index (Abl): A comparative indicator of antimicrobial potency based on inhibition zone diameter (Abl ∝ zone size).