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. 2025 Jun 18;75(3):179–186. doi: 10.1270/jsbbs.24085

Table 3. Gene families encoding enzymes involved in the rutin biosynthesis pathway and the number of genes expressed during seed formation.

Enzyme Number of loci
Detected in reference Expressed Pollen parent alleles detected
Phenylalanine ammonia-lyase (PAL) 4 4 3
Cinnamate-4-hydroxylase (C4H) 6 2 1
4-Coumarate CoA ligase (4CL) 2 2 1
Chalcone synthase (CHS) 13 5 3
Chalcone isomerase (CHI) 3 1 1
Flavanone-3-hydroxylase (F3H) 2 2 1
Flavonoid-3ʹ-hydroxylase (F3ʹH) 2 2 1
Flavonoid-3ʹ-5ʹ-hydroxylase (F3ʹ5ʹH) 16 3 3
Flavonol synthase (FLS) 2 1 0
Glycosyltransferase (GTR)a 12 (1) (1)
Total a 62 22 14

a Gene for GTR was expressed in only one plant. The total does not include GTR.