Complex genetic structure. (a) Sympatric emergence of four species. Parameters are the same as in Fig. 3 except Popt = 0.25 and S = 0.5. P̄ ≈ 0.27. (b) Genetic diversification without distinct cluster formation. Parameters are the same as in Fig. 3, except Ψij = exp[−/(2α2)], Popt = 0.7, α = 2, and initial genotype frequencies are Φ10 = 0.45, Φ−10 = 0.55. P̄ ≈ 0.71.