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. 2025 Oct 10;11:101223. doi: 10.1016/j.crfs.2025.101223

Table 2.

A comparison of research cases on the synthesis of pyrazines using different cheap carbon sources.

Carbon source type Pretreatment/processing method Key process Target product Yield Costs References
Distiller 's grains Dilute acid pretreatment; Fermentated by B. subtilis TTMP20. Optimization of detoxification conditions for distiller's grains hydrolysate. TTMP 705.27 mg/L 34-66 $/t (Gan et al., 2023; Zheng et al., 2024)
Corncob Alkali-treated; Fermentated by B. subtilis IPE5-4-UD-4. Composite mutation;Isothermal simultaneous saccharification and fermentation. Acetoin 0.46 g/g 50-30 $/t (Jia et al., 2017; Zhang, 2023)
Ignocellulosic hydrolysates Fermentated by engineered B. subtilis ZB02. The overexpression of transporter (AraE) and key enzymes (XylA and XylB) enables it to utilize glucose, xylose and arabinose simultaneously. Acetoin 11.20 g/L / Zhang et al. (2016)
Sugarcane molasses Fermentation after treatment with sulfuric acid; Soybean meal as nitrogen sources. Knockout the key genes bdhA and acoA in the acetoin catabolism pathway. TTMP 1328.95 mg/L 120 $/t (Zhou et al., 2019; Li et al., 2023)
Okara hydrolysate Batch fermentation and fed-batch fermentation; Fermentated by B. subtilis BSO3. Knockout the key genes bdhA and acoA; overexpression of the arabinose transporter-encoding gene (araE). TTMP 5.33 g/L (Batch fermentation);13.37 g/L (Fed-batch fermentation) 32-83 $/t Li et al. (2023)