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. 2025 Oct 24;2025:5351950. doi: 10.1155/vmi/5351950

Table 3.

Descriptive synthesis of outcomes resulted from melatonin supplementation of maturation media.

Outcomes Results in
CCs Oocytes Embryo development
Diameter of COCs No change [45]

Cumulus expansion Fully expanded [7]
Not expanded [7, 41, 45, 55]
Reduced expansion [55]

Live cells (%) No change [48]

DNA damage No change [47]
Reduced [43, 47]
No significant change was observed [19, 40]

Spindle disorganization Reduced significantly [50]

CG distribution Higher peripheral and cortical distribution [20, 32]

ER distribution Higher normal distribution [32]
No change [32]

Mitochondria distribution Higher normal distribution [7, 32]
No change [7, 32]

Mitochondrial activity No change [7]

MMP Higher amount [20]
No change [19]
Lower amount of MMP [19]

ATP level Higher level [32]

ROS level No change [19, 47, 51]
Lower level [7, 20, 32, 47, 48, 50, 56]
No change [19]
Lower level [49]

GSH level Higher level [20, 32]
No change [19, 48]
No change [19]

Apoptosis No change [20]
Reduced [20, 50]
Reduced [20, 48, 50]

Two pronucleus Increased [32]

Polyspermy Decreased [32]

Unfertilized oocytes Decreased [32]

Morula No change [53]

Grade 1 blastocysts No change [19]
Decreased [19]

Grade 2 blastocysts Increased [19]
No change [19]

TCN Increased [32, 34, 48, 50, 54]
No change [19, 20, 34, 44]

ICM/TE Increased [54]
No change [50]

Antioxidant-related transcripts Upregulation of CuZnSOD, MnSOD
No change in GPX4, GPX1
Downregulation of CuZnSOD [47, 57]
Upregulation of CAT, SOD1, and GPX
No change in CuZnSOD, MnSOD, and GPX4 [32, 47]
Upregulation of GPX, SOD1, SOD2, CAT, XIAP, MCL1 [19, 20, 48]

Apoptosis-related transcript Upregulation of BCL-2
Downregulation of CASP3, BAX, and SHC1 [20, 48, 57]

Development-related transcripts Upregulation of Tet1, Tet2, Tet3, GDF9, MARF1, DNMT1a
Downregulation of Dnmt1 [32, 34]
Upregulation of HSP70
No change in OCT4, NANOG, PU5F1, SLC1A, SLC2A, SLC3A, HSPB1, and KRT8 [19, 20, 48, 57]

Steroidogenic-related transcripts Upregulation of CYP11A1, CYP19A1, StAR [49]

CC-specific transcripts Upregulation of PTX3, HAS1, HAS2, LHR1, LHR2, EGFR
No change in TNFAIP6, GREM1, and PTX3 [34, 57]

Note: GSH, glutathione; TE, trophectoderm.

Abbreviations: CCs, cumulus cells; COCs, cumulus–oocyte complexes; ER, endoplasmic reticulum; ICM, inner cell mass; MMP, mitochondrial membrane potential; ROS, reactive oxygen species; TCN, total cell number.