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. 2025 Oct 31;16:1642927. doi: 10.3389/fphar.2025.1642927

TABLE 1.

Clinical trials accompanied by various biomarkers in GC.

Biomarker Type Upregulation/downregulation No. of patients Location for sampling Reference
HSPA6 Autosomal-recessive candidate protein Upregulated 166 Tissue (gene) Zhang et al. (2023b)
ANXA11 Calcium (Ca2+)-regulated phospholipid-binding proteins Upregulated 255 subjects in two stages Urine Fan et al. (2022)
CDC42 Cell division control protein 42 of the Rho family of GTPases Upregulated 255 subjects in two stages Urine Fan et al. (2022)
NAPA Protein-coding genes Upregulated 255 subjects in two stages Urine Fan et al. (2022)
FAP A type 2 membrane-bound glycoprotein Upregulated 56 normal samples and 60 GC samples Differentially expressed gene Li et al. (2022)
SERPINH1/HSP47 Serpin that serves as a human chaperone protein for collagen Upregulated 56 normal samples and 60 GC samples Differentially expressed gene Li et al. (2022)
HAMP (hepcidin antimicrobial peptide) Polypeptide hormone Upregulated 53 patients GC cells and tissues Yang et al. (2022)
Cystatin B (CSTB) A protein structure encoded by the CSTB gene Downregulated 31 healthy individuals and 31 GC (adenocarcinoma) patients Saliva Koopaie et al. (2022)
Deleted in malignant brain tumor 1 (DMBT1) A tumor-inhibiting gene Upregulated 31 healthy individuals and 31 GC (adenocarcinoma) patients Saliva Koopaie et al. (2022)
Nuclear-enriched abundant transcript 1 (NEAT1) lncRNA Downregulated 21 control samples and 111 GC samples Gene Azadeh et al. (2022)
N6-methyladenosine-related lncRNAs Internal modification in RNA Upregulated RNA sequence transcriptome data of 88 Asian GC patients
23 m6A-related genes were extracted from the TCGA database
Tissue (gene) Xu et al. (2022)
 8 circular RNAs (circRNAs) Circulating non-coding RNAs Upregulated Two independent datasets (GSE89143 and GSE83521). All expression profiling data were downloaded from the Gene Expression Omnibus (GEO) database. A pilot cohort of 28 matched GC and adjacent normal mucosal (ANM) tissues. The training cohort consisted of 92 patients with GC and 46 endoscopically negative patients (non-disease controls) enrolled at Kumamoto University, Japan, between 2010 and 2015. The second validation cohort consisted of serum specimens from 102 GC patients and 48 non-disease control subjects enrolled at the Mie University Graduate School of Medicine, Japan, between 2006 and 2017. Independent cohort of 24 GC patients from March 2017 to March 2018 at Nagoya University, Japan. Serum Roy et al. (2022)
PRTN3 A serine protease enzyme expressed mainly in neutrophil granulocytes encoded by the PRTN3 gene Upregulated 25 patients with early GC, 25 patients with progressive GC, and 25 control individuals with chronic gastritis Tissue sample Guo et al. (2023)