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. 2025 Oct 10;21(15):6580–6598. doi: 10.7150/ijbs.123492

Table 2.

Representative biomarkers of macrophage activation and metabolic dysfunction in chronic lung diseases.

Disease Biomarker Source Macrophage-derived? Clinical Relevance
Asthma Arginase-1 BALF macrophage Yes (M2 macrophage) Promotes collagen synthesis, airway fibrosis
YKL-40 (CHI3L1) 67 Serum/BALF Yes (macrophages, others) Correlates with asthma severity and airway remodeling
FeNO 160 Breath Partially (epithelium via iNOS; modulated by Arg1) Elevated in allergic (type 2) asthma; indicates airway eosinophilic inflammation; influenced by macrophage arginase activity
LTB4 Sputum/BALF Yes (macrophages, neutrophils) Increased in asthmatic airways; contributes to bronchoconstriction and inflammation
15-HETE & Lipoxin A4 BALF Yes (macrophages) Reduced in asthma; anti-inflammatory and pro-resolving lipid mediators
CCL17 (TARC) BALF Yes Macrophage-secreted chemokine; eosinophil/T cell recruitment
Periostin Serum No Biomarker of type 2 inflammation
COPD CXCL8 (IL-8) Sputum Yes (macrophages, epithelium) Markedly elevated in COPD sputum; drives neutrophil recruitment
MMP-9, MMP-12 BALF/Sputum Yes (macrophages, neutrophils) Promote elastin degradation and emphysema progression
3-Nitrotyrosine Sputum cells Yes (macrophages) Reflects peroxynitrite (ONOO-)-mediated oxidative stress
Hemosiderin 161 BALF macrophages Yes (indicative) Indicates iron overload and oxidative injury
SP-A, SP-D 162 Serum No General marker of lung injury and exacerbations
IPF CCL18/PARC 163, 164 Serum, BALF Yes (M2 macrophages) Reflects profibrotic M2 activation; predicts poor survival and faster disease progression
YKL-40 (CHI3L1) Serum, BALF Yes (macrophages, others) Associated with fibrosis and matrix remodeling
MMP-7 165 Serum Partially (epithelium, macrophages) Indicates active epithelial injury and remodeling
KL-6 (MUC1) 164, 166, 167 Serum No (alveolar type Ⅱ cells) Widely used diagnostic and prognostic biomarker for ILDs
SP-D 164, 168 Serum No Reflect epithelial injury and surfactant disturbance
Reduced itaconate (ACOD1 product) 114 BALF macrophages Yes Loss of anti-inflammatory metabolite regulation; linked to persistent activation
CD44 169 BALF/lung tissue Partially (macrophages, fibroblasts) Regulates macrophage-fibroblast crosstalk; associated with fibrosis progression
CX3CL1 (fractalkine) 170 Serum, BALF Yes (Endothelium, macrophages) Potential diagnostic biomarker; correlates with IPF progression
S100A8/A9/A12 (S100 family) 171 Serum, BALF Partially (macrophages, neutrophils) Associated with inflammation, oxidative stress, and fibrogenesis

Abbreviations: BALF, bronchoalveolar lavage fluid; FeNO, fractional exhaled nitric oxide; LTB4, leukotriene B4; 15-HETE, 15-hydroxyeicosatetraenoic acid; CCL17, chemokine (C-C motif) ligand 17; TARC, thymus and activation-regulated chemokine; MMP, matrix metalloproteinase; SP, surfactant protein; CCL18/PARC, pulmonary and activation-regulated chemokine; KL-6, Krebs von den Lungen-6, a mucin 1 glycoprotein; ILDs, interstitial lung diseases; ACOD1, aconitate decarboxylase 1.