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. 2025 Nov 28;16:1701101. doi: 10.3389/fmicb.2025.1701101

Table 1.

Gut microbiota alterations in hepatocellular carcinoma patients compared to healthy individuals.

Category Microbiota/parameter Taxonomic level Change in HCC Clinical/pathological significance Refs
Community Overall gut microbial diversity Ecosystem ↓Decreased Dysbiosis, reduced ecosystem stability, and resilience Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Phylum (↑) Proteobacteria Phylum ↑ Increased Liver tissue enrichment; associated with inflammation Yu and Schwabe (2017)
Phylum (↑) Actinobacteria Phylum ↑ Increased Liver tissue enrichment; pro-inflammatory potential Yu and Schwabe (2017)
Phylum Bacteroidetes, Firmicutes Phylum Altered* Metabolic and immune dysregulation Yu and Schwabe (2017)
Taxa (↑) Rhizobiaceae Family ↑ Increased Microbial signature associated with HCC Dossa et al. (2016)
Taxa (↑) Agrobacterium Genus ↑ Increased Microbial signature associated with HCC Dossa et al. (2016)
Taxa (↑) Parabacteroides Genus ↑ Increased Linked to hepatic inflammation and fibrosis Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↑) Clostridium Genus ↑ Increased Associated with HCC development and progression Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↑) Gemmiger Genus ↑ Increased Promotes inflammatory responses Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↑) Phascolarctobacterium Genus ↑ Increased Contributes to hepatic pathology Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↑) Enterococcus Genus ↑ Increased Pathogenic potential, inflammation driver Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↑) Streptococcus maltophilia Species ↑ Increased Strongly associated with poor prognosis Dossa et al. (2016)
Taxa (↓) Pseudomonas Genus ↓ Decreased Loss of potentially protective bacteria Dossa et al. (2016)
Taxa (↓) Verrucomicrobiaceae Family ↓ Decreased Impaired intestinal barrier function Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Bifidobacteriaceae Family ↓ Decreased Reduced protective and anti-inflammatory effects Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Akkermansia Genus ↓ Decreased Compromised mucus layer integrity Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Alistipes Genus ↓ Decreased Immune dysregulation Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Dialister Genus ↓ Decreased Loss of barrier-protective functions Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Collinsella Genus ↓ Decreased Altered metabolic homeostasis Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Taxa (↓) Adlercreutzia Genus ↓ Decreased Reduced beneficial metabolite production Rajapakse et al. (2023), Behary et al. (2021a), and Zhang et al. (2021)
Metabolite TMAO (Trimethylamine N-oxide) Microbial metabolite ↑ Increased (serum) Positively correlated with HCC risk; promotes tumor progression Zhou et al. (2024) and Liu et al. (2018)
Metabolite SCFAs (Short-chain fatty acids) Microbial metabolite ↓ Decreased Loss of anti-inflammatory, barrier-protective, and anti-tumor effects Liu J. et al. (2023), Guo et al. (2013), and Gao et al. (2021)
Biomarker LPS antibodies Immune marker ↑ Increased (serum) Indicates bacterial translocation and barrier dysfunction Yang et al. (2019)
Biomarker Flagellin antibodies Immune marker ↑ Increased (serum) Reflects gut-liver axis dysregulation and systemic inflammation Yang et al. (2019)

Changes indicate alterations in HCC patients compared to healthy individuals. ↑ = increased abundance/levels; ↓ = decreased abundance/levels. *Bacteroidetes and Firmicutes are noted as altered in HCC patient liver tissue, but specific directional changes require further clarification. TMAO = Trimethylamine N-oxide; SCFAs = Short-chain fatty acids; LPS = Lipopolysaccharide. Refs = Reference numbers from manuscript. Color coding: Red = increased taxa; Green = decreased taxa; Yellow = metabolites; Blue = biomarkers.