Table 2. Qualitative morphological traits.

Diagnostic characters used for comparison of Pristimantis paganus sp. nov. and its nominal sister species in the Cryptomantis subgenus.

Species	SVL (mm)		Snout in dorsal view	Snout in profile view	Disc shape	Dermal fold	Upper eyelid tubercles	Lateral cutaneous ridges on hands	Ulnar tubercles	Tubercles in heels	Tubercles on external border of the tarsus	Altitudinal range (m.a.s.l.)
	Female	Male
P. paganus sp. nov.	24.84–29.8	17.20–21.5	Subacuminate	Rounded	Expanded	Defined and discontinuous dorsolateral dermal folds on the dorsum in females, defined continuous in males	Present, 3 to 4 rounded tubercles surrounded by several small ones.	Present	Absent	Subconical tubercle	Two flattened tubers	2,157–2,495
P. bicantus	17.0–21.7	12.0–15.8	Rounded	Rounded	Slightly expanded	Absent	Present, low tubers	Absent	Absent	Absent	Absent	2,000–2,300
P. burtoniorum	20.8–27.0	16.6–17.4	Subacuminate	Subacuminate	Expanded discs	Absent	Present, three to four large subconical tubers	Present	Small, conical	Small conical tuber	Present, round-shaped	2,940–2,970
P. ganonotus	–	14.0–15.2	Acuminate	Protruding	Expanded	Absent	Absent	Absent	Indistinct	Nonconical	Absent	2,940–2,970
P. marcoreyesi	22.8–30.8	14.3–22.2	Subacuminate	Angular	Slightly wider than the digits	Present, weakly defined	Present, two or more flattened to subconical tubercles present	Present	Small, flattened	Presents, rounded	Present, weakly defined	2,500–3,100
P. nelsongalloi	12.0–17.0	18.5–21.7	Subacuminate	Rounded	Lanceolate discs	Dorsolateral folds thin anteriorly and discontinuous posteriorly	Present, four or five small supraoculars, flat and low	Absent	Present, small	One subconical and several small tubers	Absent	1,627–1,800
P. prolatus	20.8–24.1	13.7–18.4	Acuminate	Truncated	Expanded	“H” shaped	Present, one conical	Absent	Absent	Present, conical	Present	1,140–1,933
P. sacharuna	18.5–19.5	–	Subacuminate	Slightly rounded	Slightly wider than the digits, expanded, rounded	W-shaped scapular fold followed by fine scapular fold in shaped of an inverted “V”	Present, one subconical	Present	Present, subconical	Subconical tubercle	Two to 3 subconical tubers	2,200
P. tungurahua	24.4–27.9	17.1–20.8	Subacuminate	Protuberant	Rounds, expanded	W-shape very evident in males, not very evident in females	Present, one or two subconical tubercles	Absent	Present, low	Present, conical	Present, conical	2,500–2,750