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. Author manuscript; available in PMC: 2026 Jan 19.
Published in final edited form as: Nat Protoc. 2024 Feb 2;19(5):1311–1347. doi: 10.1038/s41596-023-00948-y

Table 2 |.

Effects of common cell metabolism-perturbing agents on subcellular lactate metabolism

Pathway Agent Mechanism(s) of action Stock solution Final solution Cytosolic changes Nuclear changes Mitochondrial changes Extracellular changes
Glycolysis 3-BrPA39 HK II 100 mM in HBSS-G 500 μM Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease
2-DG HK II 500 mM in HBSS-G 50 mM Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease
Oxamate39 LDH 400 mM in HBSS-G 10 mM Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease
IAA GAPDH 10 mM in HBSS-G 500 μM Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease
Oxalate PK 5 mM in HBSS-G 500 μM Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease Free [lactate] decrease
UK5099 MPC 10 mM in DMSO 5–10 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
TCA Devimistat PDH/aKGDH 100 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] decrease Free [lactate] increase
ETC Rotenone39 Complex I 30 mM in DMSO 10 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Oligomycin A39 Complex V 5 mM in DMSO 5 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
pH Regulation AZD-3965 (ref. 39) MCT1 10 mM in DMSO 2 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] decrease
AR-C155858 (ref. 39) MCT1/2 10 mM in DMSO 2 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] decrease
Fatty acid metabolism SB-204990 (ref. 39) ACLY 30 mM in DMSO 50 μM Free [lactate] increase Free [lactate] increase Free [lactate] decrease Free [lactate] increase
BMS-303141 (ref. 39) ACLY 30 mM in DMSO 10 μM Free [lactate] increase Free [lactate] increase Free [lactate] decrease Free [lactate] increase
Perhexiline maleate CPT1 30 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Etomoxir CPT1 30 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
JZL195 MAGL 30 mM in DMSO 30 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
KML29 MAGL 15 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
mTOR Rapamycin39 mTOR 10 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Metabolites Octyl-(R)-2-HG39 ATP synthase/mTOR 100 mM in DMSO 500 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Octyl-(S)-2-HG39 ATP synthase/mTOR 100 mM in DMSO 500 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Ca2+ Regulation Ionomycin39 Calcium ionophore 10 mM in DMSO 2 μM Free [lactate] increase Free [lactate] increase Free [lactate] decrease Free [lactate] increase
Medical/health Drug CoQ39 Essential Cofactor of ETC 10 mM in ethanol 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
MitoQ39 Mitochondri-targeted antioxidant 10 mM in DMSO 10 μM Free [lactate] increase Free [lactate] increase Free [lactate] increase Free [lactate] increase
Troglitazone39 Antidiabetes medicine 100 mM in DMSO 100 μM Free [lactate] increase Free [lactate] increase Free [lactate] decrease Free [lactate] decrease

Bold indicates an increase in the concentration of free Lactate ('free [Lactate]'). 3-BrPA, 3-bromopyruvic acid; HK II, hexokinase II; IAA, iodoacetamide; GAPDH, gLyceraLdehyde-3- phosphate dehydrogenase; PK, pyruvate kinase; MPC, mitochondrial pyruvate carrier; TCA, tricarboxylic acid; ETC, electron transport chain; PDH, pyruvate dehydrogenase; α-KGDH, α-ketoglutarate dehydrogenase; ATMA, antimycin A; MCT1/2, monocarboxylate transporter 1/2; ACLY, ATP citrate lyase; CPT1, carnitine palmitoyltransferase 1; MAGL, monoacylglycerol lipase; mTOR, mammalian target of rapamycin; 2-HG, 2-hydroxyglutarate; CoQ, Coenzyme Q10; MitoQ, Mitoquinone.