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Probability 
That a Founder with f Copies of Marker Allele B1 Carries i Copies of a Nearby Disease Allele
| f |  |
 |
 |
 |
 a
|
| 2 | q21 | 2p1q1 | p21 | 2c1p1+ep21b | 1 |
| 1 | q1q2 | p1q2+q1p2 | p1p2 | c1(p1+p2)+ep1p2 | λc |
| 0 | q22 | 2p2q2 | p22 | 2c1p2+ep22 | 0 |
, if and only if Δ≠0, where Δ=p1-p2 is the standardized disequilibrium coefficient.
e=1-2c1.
.
