Movement rates of butterfly species between connected and isolated patches. J. coenia (a) and E. claudia (b) both moved between connected patches more often than between isolated patches (J. coenia: F1,22 = 12.0, P = 0.001; E. claudia: F1,22 = 5.3, P = 0.016). There was no indication of a drift-fence effect (comparing winged to rectangular patches, both P > 0.7). Data in both panels are means ± 1 SE for proportion of individuals marked in the central patch and recaptured in connected, winged, and rectangular peripheral patches.