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(a) Signal flows in the radial lines of the structural model. Blue arrows—synaptic flux in layer 4. Red arrows—synaptic flux in layers 5,6 and 2,3. Markov blankets form at the 2,3 versus 4 junction. (b) Formation of circumferential interareal connections. Cortical areas bilaterally coupled by cortico-cortical fibers (gray arrows) link multi-columnar systems within each area.
