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. Author manuscript; available in PMC: 2026 Aug 4.
Published in final edited form as: Handb Clin Neurol. 2018;148:441–464. doi: 10.1016/B978-0-444-64076-5.00029-6

Table 29.1.

PRNP missense mutations

PRNP mutation Codon 129 polymorphism # of cases in literatureb Clinical phenotypes Age at onset (range)f (years) Disease duration (months or years) Positive family historyc CSF marker sensitivity
EEG PSWC MRI c/w JCDd Neuropathology Neuropathology phenotype References
14-3-3 Total taug
P84S MV 1 Cog (1 year) → paranoia and RPD (60) (14) months 0% (0/1) N/A 0% (0/1) 0% (0/1) Multicentric
PrP-Plqs w/o NFT
No V
GSS Jones et al. (2014)
S97N Cis M 1 AD (72) N/A 0% (0/1) N/A N/A 0% (0/1) N/A N/A Zheng et al. (2008)
P102L MM/MV (most cis M) ~221 Early Cb w/ late D
Some are RPD
LE areflexia common
(27–66) (7–132) months 84–100% (15–20% ) 20% 0–35% 25–30% Multicentric
PrP-Plqs
GSS Webb et al. (2008); Higuma et al. (2013); Krasnianski et al. (2016)
P102La Cis M 13 44 ± 12 (24–57) 44 ± 13 months (28–60) FHx score
0 (n = 1)
1 (n = 2)
2 (n = 6)
0% (0/3) 0% 0% (0/3) 33.3% (1/3) Takada et al. (2017))
P105L MV 13 D w/ spastic paraparesis
Cb Atx
Pscyh Sxs sometimes
Mean 44 ± 10 (2nd to 7th decades) 111 ± 82 months 37% 0% 0% 0% 14% PrP-Plqs
diff PrP (deep CLs)
GSS Higuma et al. (2013)
P105La N/A 1 (9) N/A FHx Score
0 (n = 1)
N/A N/A 0% (0/1) 0% (0/1) N/A Takada et al. (2017)
P105T Cis M 13 Usually RPD;
Cb Atx freq
(13–41) (2-5) years 100% (2/2) 0% N/A 0% (0/3) 25% (1/4) V, PrP-S (all CLs) Unicentric PrP-Plqs
(deep CLs)
JCD Rogaeva et al. (2006);Polymenidou et al. (2011)
P105S MV (cis V) 1 Aphasia, frontal-type behavioral changes, D, late Park (30) (10) years 0% (0/1) N/A N/A 0% (0/1) 100% (1/1) Multicentric PrP-Plqs (HP), punctate aggregates (Cb), V (Pu) Atypic GSS Tunnell et al. (2008)
G114V (MM/MV) 1 RPD
Onset: Psych Sxs, D, Park, Pyram signs, myoclonus
GTCs in some Absent or mild Cb signs
(18–75) (1–4) years 75% (3 in 4 probands) e 0% 0% (0/8) 42.9% (3/7) predom BG Mod V, G, NL. PrP-S, type 1 PrPSc (predom monoglycos) JCD Rodriguez et al. (2005); Ye et al. (2008); Beck et al., (2010);Liu et al. (2010)
A117V Cis V 33 Variable
Progressive D w/o Atx
LMN SYN w/ D and Atx
(20–64) (1–11 years) 100% (4/4) N/A N/A N/A 0% (0/2) Ab PrP-Plqs, F V, NL, G GSS Hsiao et al. (1991); Mastrianni et al.(1995); Kong et al. (2004)
A117V* Cis V 6 34 ± 14 (14–49) 46 ± 21 months
(27–78)
FHx score
0 (n = 1)
2 (n = 2)
0% 0% 0% (0/3) 0% (0/5) Takada et al. (2017)
G131V MM/MV (cis M) 3 D w/ behavioral changes and late Atx
Park
(36–42) (9–16) years 50% (1/2) N/A N/A 0% (0/1) 0% (0/1) PrP-Plqs, NFT (AH, ERC), No V GSS Panegyres et al. (2001); Jansen et al. (2012)
S132I MM 2 RPD (62) (18) months 100% (1/1) N/A N/A N/A N/A Diff unicentric and multicentric PrP-PLs
(neocortex, BG, Cb)
Min V
GSS Hilton et al. (2009)
A133V MM 2 PSP-like, RPD (62) (4) months 0% (0/1) 0% 0% (0/1) 0% (0/1) Diff V, G, NL multic PrP-Plqs in the (Mol), PrP-S (Th) Atypic GSS Rowe et al. (2007)
R148H (MV/MM) 3 JCD (62–82) (6–18) months 0%
(0 in 2)e
50% (1/2) 100% (1/1) 50% (1/2) 100%
(1/1) predom BG
129MM - similar to sJCDMM1. V, G, NL (deeper CLs). PrP-S
PrPSc type 1 129MV - similar to
sJCDMV2 V predom in CLs
V and VI, Kuru plaques in Cb and
WM, PrP-S PrPSc type 2 (predom monoglycos)
JCD Krebs et al. (2005); Pastore et al.(2005)
R148H* MM 1 (53) (2) months FHx score 0 (n = 1) N/A N/A N/A 100% (1/1) Takada et al. (2017)
D167G MM 1 JCD N/A N/A N/A N/A N/A N/A N/A sJCD PrP type 1 JCD Bishop et al. (2009)
D167N MM 1 RPD w/ Park and Pyram signs (33) (2) years 0% (0/1)e N/A N/A 0% (0/1) 0% (0/1) N/A N/A Beck et al. (2010)
V176G VV 1 RPD w/ behavioral changes, Cb Atx, Pyram signs and myoclonus (61) (7) months 0% (0/1) 100% (1/1) 100% (1/1) 0% (0/1) 0% (0/1) Multic PrP-Plqs w/ prominent tau GSS Simpson et al. (2013)
D178N-129V MV/VV (cis V) 209a Progressive Cog decline, Cb Sxs, myoclonus, EP Sxs Mean 46 (26–56) Mean 23 months
(7–60)
100% (12/12) N/A N/A N/A N/A Similar to sJCD VV1 JCD Brown et al. (1992); Goldfarb et al., (1992); Kong et al. (2004)
D178N-129Va Cis V 8 45 ± 6 (37–52) (18–21) months FHx score 2 (n = 4) 50% (1/2) 0% (0/1) 0% (0/2) 100% (2/2) Takada et al. (2017)
D178N-129M Cis M 106 Insomnia Sympathetic overactivity Mean 52 (~20–76) Median 12.6 (4–40) months 60–88% 14.4% 8% 2.1% 16.5% Deg of Th (md and av nu) and inf olivary nu, little to no Vor PrPSc dep FFI Reder et al. (1995); Collins et al. (2001); Kovacs et al. (2005); Zarranz et al. (2005); Sano et al. (2013); Krasnianski et al. (2016)
V180I Cis M 225 JCD phenotype, but w/ slower prog Mean 77 Mean 25 months 0.7–6% 70%; 78.5% 11% 99% V PrP-S JCD Kong et al. (2004); Higuma et al. (2013)
V180I* Cis M 2 (84) (21) months FHx score
1 (n = 1)
2 (n = 1)
0% (0/1) 0% (0/1) 100% (1/1) Takada et al. (2017)
T183A Cis M 3 bvFTD, AD 45 ± 4 (42–49) 4 ± 2 years
(2–9)
100% (2/ 2) N/A N/A 0% (0/7) 0% (0/2) V & NL (CLs IV, V, VI), PrP (Cb, Pu) Small Plq-like PrP, Predom monoglycos PrPSC JCD Nitrini et al. (1997); Grasbon-Frodl et al. (2004)
H187R MM/MV/VV 7 Early Cog and
behavioral Sxs w/late Cb Atx
Early Cb Atx and D after a few years
Case w/ Pscyh Sxs in adolescence
(20–53) (3–19) years 100% (4/4) 0% (0/2) 0% (0/4) 0% (0/4) G multicentric PrP-Plqs in some cases. Curly PrP-G GSS Cervenakova et al. (1999);
Butefisch et al. (2000); Hall et al. (2005); Colucci et al. (2006)
H187R* Cis M 4 (30–41) (12–13) years FHx score 2 (n = 1) 0% (0/1) 0% (0/1) 0% (0/1) 0% (0/1) Takada et al. (2017)
T188R MV/VVV (cis V) 12 JCD (55–66) (14–16) months 0% (0/1)e 50% (1/2) 100% (1/1) 50% (1/2) 50% (1/2) V, NL, A PrP-S and plaque-like PrP, type 1 PrPSc JCD Roeber et al. (2008); Tartaglia et al.(2010)
T188K MM/MV (cis M) 3 JCD Median 58 (39–76) (2–13) months 8–37%e 69% 12% 69% SE PrP-S JCD Roeber et al. (2008); Chen et al. (2013); Shi et al. (2015)
T188A MM 1 JCD (82) (4) months 0% (0/1) 100% (1/1) 100% (1/1) 100% (1/1) 0% (0/1) Sev G, V, Mod NL (predom in OLs) PrP-Neg JCD Collins et al. (2001)
T193I MM JCD (70) (10) months 0% (0/1) 100% (1/1) 100% (1/1) 100% (1/1) 0% (0/1) N/A N/A Kotta et al. (2006)
E196K N/A 13 JCD (64–69) (10–13) months 100% (1/1) N/A N/A 0% (0/1) N/A N/A N/A Peoc’h et al. (2000)
F198S MV/VV 5 Cb Atx and D freq Park (40–71) years Mean 5 years (2–12) 100% (3/3) N/A N/A N/A 0% (0/1) Uni- and multicentric PrP-Plqs GSS Farlow et al. (1989); Dlouhy et al. (1992); Ghetti et al. (1995); Kong et al. (2004)
F198S* Cis V 5 55 ± 8 (46–66) 67 ± 23 months
(34–84)
FHx score
1 (n = 2)
2 (n = 1)
33.3% (1/3) 100% (1/1) 0% (0/4) 0% (0/3) Takada et al. (2017)
F198V MM 1 D w/ visual
hallucinations, myoclonus and Park
Clinical dx of early-onset AD
(56) (4) years N/A N/A N/A 0% (0/1) 0% (0/1) N/A N/A Zheng et al. (2008)
E200K MM/MV/VV 571 Similar to sJCD Peripheral
neuropathy and supranuclear gaze palsy in some
Mean 60 (33–84) (1–18) months 50% 85–100% 80–100% 42–85% 50–88% Usually sJCD MM1 PrPSc types 1 and 2 JCD Spudich et al. (1995); Meiner et al. (1997); Kovacs et al., (2005, 2011); Krasnianski et al. (2016)
E200K* Cis M (n = 16) and cis V (n = 1) 34 60 ± 13 (36–84) 11 ± 17
(1–78)
FHx score
0 (n = 2)
1 (n = 10)
3 (n = 12)
57.1% (4/7) 37.5% (3/8) 88.9% (16/18) Takada et al. (2017)
E200G MV (cis V) 1 RPD, Cb Atx, Park ↓sensation in LEs (57) (30) months 0% (0/1) 0% (0/1) 100% (1/1) 0% (0/1) 100% (1/1) SE w/ type 2 PrPSc JCD Kim et al. (2013)
D202G MV (cis V) 1 Slowly progressive D w/ Cb Atx Later Pyram and EP signs (55) (16) years 100% (1/1) 100% (1/1) 0% (0/1) 0% (0/1) 0% (0/1) N/A N/A Heinemann et al. (2008)
D202N VV 1 D (AD) w/ Cb Atx (73) (6) years N/A N/A N/A N/A N/A PrP-Plqs, NFT GSS Piccardo et al. (1998)
V203I N/A 17 JCD (69) (1) month 0% (0/1) N/A N/A 100% (1/1) N/A N/A N/A Peoc’h (2000)
R208H MM/VV 15 D w/ behav changes Park and Pyram
signs freq Report of a PSP-like phenotype
(58–63) (3–16) months 20% (1/5) 50% (4/8) 57.1% (4/7) 25% (2/8) SE, PrP-S (perineuronal perivacuolar) type 1 PrP JCD Capellari et al. (2005); Roeber et al. (2005); Matej et al. (2012); Vita et al. (2013); Shi et al. (2015)
V210I Cis M 247 JCD Mean 59 (39–82) Median 5 (2–20) months 12–31% 90–100% 100% 44–80% 15–33% Similar to sJCD MM1 JCD Kong et al. (2004);Kovacs et al. (2005);Breithaupt et al. (2013);Krasnianski et al. (2016)
V210I* Cis M 3 57 ± 15 (47–74) (1) month FHx score
0 (n = 2)
2 (n = 1)
100% (1/1) 33.3% (1/3) 100% (2/2) Takada et al. (2017)
E211Q MM 11 JCD (42–81) (6–32) months 100% (2/2) N/A N/A 100% (4/4) N/A V, G Mi
PrP-S types 1 and 2 PrPSc
JCD Peoc’h et al. (2000, 2012); Ladogana et al. (2001)
E211D VV 1 Cb Atx, and late D (53–68) (3 –13) years 50% (1/2) N/A N/A 0% (0/2) 0% (0/2) Multicentric PrP-Plqs
Dystrophic neurites and NFT
GSS Peoc’h et al. (2000, 2012)
Q212P MM 2 Cb Atx w/o D Dx of olivoponto Cb degeneration (60) (8) years N/A N/A N/A N/A N/A Mod PrP Mi, PrP-Plqs GSS Piccardo et al. (1998)
I215V MM 1 JCD (55–76) (12–15) months 0% (0/2) (1/3) 100% (2/2) 50% (1/2) NL, G, V, PrP-Neg JCD
Q217R VV/MV (cis V) 3 D w/ Cb Atx Cog decline, stereotypical behav Late Park and apraxia. Clinical dx of bvFTD and CBS (45–66) (5–13) years 100% (3/3) N/A N/A 0% (0/1) 0% (0/1) Uni- and multicentric PrP-Plqs, NFT (neocortex) GSS Hsiao et al. (1992); Piccardo et al. (1998); Woulfe et al. (2005); Munoz-Nieto et al. (2013)
Y218N VV 1 Atypic D w/AD and (54–61) bvFTD features Early language and executive impairment No Atx (6) years 100% (1/1) N/A N/A 0% (0/2) 0% (0/2) Uni- and multicentric PrP-Plqs, NFT w/ hyperP tau GSS Alzualde et al. (2010)
A224V VV (cis V) 1 RPD (48) (32) months 0% (0/1) e 100% (1/1) N/A 100% (1/1) Diff V w/ PrPSc type 1 JCD Watts et al. (2015)
M232R MM 63 Similar to sJCD, some w/ slower prog Mean 64 (15–81) Mean 8 (0–32) months ~0% 55–75% 55–93% 20–100% 85% sJCD MM1 JCD Shiga et al. (2007); Zheng et al. (2008);
Nozaki et al. (2010); Higuma et al. (2013)
M232T MV Cb Atx, spastic paraparesis and D N/A (6) years 0% (0/1) N/A N/A N/A N/A Multicentric PrP-Plqs GSS Bratosiewicz et al. (2000)
P238S N/A JCD N/A N/A N/A N/A N/A N/A N/A N/A N/A Windl et al. (1999)
a

Including D178N-129V and D178N-129M.

b

By nine prion disease (PrD) surveillance centers, according to Minikel et al. (2016).

c

Positive family history of dementia with similar clinical features (as of the proband) or PrD. For UCSF FHx (family history) score scale: 0 when there was no positive family medical history suspicious for or known PrD; 1 when there was at least one first-degree relative with dementia, encephalopathy, or movement disorder; or 2 in patients who were part of families with known PRNP mutations, or had positive history for clinical or path-proven PrDs.

d

According to most commonly used European 2009 and UCSF 2011 criteria (Zerr et al., 2009; Vitali et al., 2011).

e

There is evidence of incomplete penetrance, as asymptomatic older carriers also were identified.

f

Data on age at onset and duration of disease are shown as mean ± sd (range), unless otherwise indicated.

g

positive if > total tau 1200 pg/mL.

If there are differences between data published in the literature from the more recently published University of California, San Francisco (UCSF) cohort, this information is provided in the table separately for that mutation.

Ab, abundant; AD, Alzheimer-type dementia; AH, Ammon horn; atypic, atypical; Atx, ataxia; av, anteroventral; BG, basal ganglia; bvFTD, behavioral variant frontotemporal dementia; Cb, cerebellum; CBS, corticobasal syndrome; CLs, cortical layers; Cog, cognitive; CSF, cerebrospinal fluid; c/w, consistent with; D, dementia; deg, degeneration; dep, deposition; diff, diffuse; dx, diagnosis; EEG, electroencephalogram; EP, extrapyramidal; ERC, entorhinal cortex; FFI, fatal familial insomnia; FHx, family history; F, focal; freq, frequent; G, gliosis; GSS, Gerstmann–Sträussler–Scheinker; GTC, generalized tonic-clonic seizures; HP, hippocampus; HyperP, hyperphosphorylated; inf, inferior; JCD, Jakob–Creutzfeldt disease; LE, lower-extremities; LMN, lower motor neuron; md, mediodorsal; Mi, mild; Min, minimal; Mod, moderate; Mol, molecular layer of the cerebellum; monoglycos, monoglycosylated; MRI, magnetic resonance imaging; N/A, not available; NFT, neurofibrillary tangles; NL, neuronal loss; nu, nuclei; OLs, occipital lobes; Park, parkinsonism; predom, predominant; prog, progression; PrP-G, granular PrP deposits; PrP-Neg, negative PrP staining; PrP-Plqs, PrP-amyloid plaques; PrP-S, synaptic PrP deposits; PSP, progressive supranuclear palsy; PSWC, periodic sharp-wave complexes; Pu, putamen; Pyram, pyramidal; Psych, psychiatric; RPD, rapidly progressive dementia; SE, spongiform (vacuolated) encephalopathy; Sev, severe; sJCD, sporadic Jakob–Creutzfeldt disease; SYN, syndrome; Th, thalamus; Sxs, symptoms; V, vacuolation; w/, with; WM, white matter; w/o, without.