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. 2026 Jul 21;20:1835506. doi: 10.3389/fnins.2026.1835506

Table 2.

Representative lipophilic cations compounds: chemical features and neurodegenerative-disease fit.

Compound (TPP+ conjugate) Cargo class/core chemistry Primary mitochondrial action Chemical advantages Chemical liabilities Best-fit NDD context (conceptual) Key translational bottlenecks
MitoQ (Murphy and Smith, 2007; Smith et al., 2012) Quinone (ubiquinone/ubiquinol cycling) Redox cycling; lipid peroxidation suppression Sustained activity via ETC-mediated reduction; strong IMM association Context-dependent pro-oxidant risk at high accumulation; ΔΨm dependence AD/PD where ROS and ETC stress are early features BBB exposure; chronic dosing safety; stage dependence
SkQ1 (Jiang et al., 2018; Skulachev et al., 2009) Plastoquinone derivative Cardiolipin-associated antioxidant protection High membrane/lipid targeting; cristae stabilization potential High lipophilicity may increase nonspecific membrane perturbation AD/aging-related synaptic mitochondrial fragility; inflammation-linked injury Long-term safety; distribution control (Zielonka et al., 2017)
MitoTEMPO/MitoTEMPOL (Dikalov and Harrison, 2014; Trnka et al., 2008) Nitroxide radical (SOD mimetic) Catalytic ROS scavenging; oxidative stress attenuation Catalytic-like ROS handling; strong fit for dopaminergic oxidative stress Radical chemistry can have off-target reactivity; PK variability PD (dopamine/iron-driven ROS burden) Exposure/retention in brain; dose-limited toxicity
MitoVitE (McCormick et al., 2016; Plecitá-Hlavatá et al., 2009; Smith et al., 2003) α-Tocopherol (lipid antioxidant) Inhibits lipid peroxidation in membranes Strong membrane protection; synergy with ROS reduction Very lipophilic → nonspecific membrane binding; accumulation toxicity risk AD where lipid peroxidation/cardiolipin damage contributes to synapse loss Safety window; formulation challenges
MitoGSH (Kezic et al., 2016; Liu Y. et al., 2023; Marí et al., 2020) Glutathione-linked thiol system Enhances endogenous redox buffering Aligns with native antioxidant networks; complements catalytic scavengers Thiol instability/metabolism; limited persistence Broad NDDs with redox imbalance (AD/PD/ALS) Bioavailability; stability; sustained efficacy
MitoNAC (Cheng et al., 2023; Vilas-Boas et al., 2023) N-acetylcysteine-linked thiol Supports GSH synthesis; redox buffering Simple chemistry; boosts endogenous defense Similar thiol limitations; dosing may be high Broad NDDs; potentially ALS where oxidative stress is prominent BBB penetration; chronic regimen practicality
Mito-metformin (Ay et al., 2024; Loan et al., 2024) Biguanide (metabolic modulator) Bioenergetics/biogenesis signalling modulation Repurposing logic; targets metabolic component of NDDs Risk of mitochondrial over-inhibition if excessive; ΔΨm dependence PD/aging-related mitochondrial biogenesis deficits Mechanism complexity; tolerability and dose optimization
Rhodamine derivatives (Chauhan et al., 2022; Silachev et al., 2015) Delocalized aromatic cation dye Mitochondrial imaging; carrier-assisted antioxidant delivery Strong membrane permeability; intrinsic fluorescence; tunable chemistry Phototoxicity risk; limited therapeutic potency alone AD/PD mitochondrial dysfunction mapping; theranostics Stability; toxicity; limited drug payload capacity
Dequalinium (DQA)/DQAsomes (Mursaleen et al., 2023; Yue Zhang et al., 2021) Dicationic amphiphile vesicle-forming system Gene and small-molecule mitochondrial delivery Self-assembly; macromolecule delivery potential; dual targeting Complex PK; cellular toxicity at high dose Emerging for neurodegeneration gene modulation; mitochondrial repair Scale-up; BBB delivery; safety profile
Pyridinium-based cations (Xu et al., 2022; Zielonka et al., 2017) Aromatic heterocycle cation ROS sensing; mitochondrial phototherapy; cargo conjugation Electronic tunability; optical activity; modular synthesis Potential off-target reactivity; limited clinical experience AD imaging; anti-aggregation strategies Translational validation; pharmacokinetics
Cyanine dyes (mitochondria-targeted) (Kim J. et al., 2023; Yan et al., 2024) Extended conjugated cationic dye Imaging; photothermal/photodynamic modulation Strong mitochondrial accumulation; BBB penetration; theranostics Phototoxicity; metabolic instability Neuroinflammation imaging; mitochondrial dysfunction monitoring Safety; light-based therapy limitations
Berberine derivatives (cationic alkaloids)(Aryal et al., 2022; Yuan et al., 2019) Planar bioactive cation scaffold Modulation of mitochondrial metabolism and ROS Natural product origin; BBB penetration; multi-target activity Off-target signaling; variable mitochondrial specificity AD/PD metabolic dysfunction; neuroinflammation Dose optimization; specificity