Organization of the operons hya, hyb, hyc, and hyp. Homologous genes have the same type of shading. Black boxes, large subunit genes; gray boxes, small subunit genes; hatched boxes, genes for the specific endopeptidases; crosshatched boxes, genes for cytochrome b; vertically striped boxes, genes for HypC and its homolog HybG; horizontally striped boxes, genes for HypA and its homolog HybF.