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Production of holomycin in SA medium by the wild-type S. clavuligerus strain ATCC 27064 (○) and the following clavulanic acid-nonproducing strains: (A) S. clavuligerus ccaR::aph (•); (B) S. clavuligerus claR::aph (▪) and S. clavuligerus car::aph (□); (C) S. clavuligerus cyp::aph (▵), S. clavuligerus ORF12::aph (▴), and S. clavuligerus ORF15::aph (⋄).
