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Formation of holomycin by the disrupted S. clavuligerus mutant ORF15::aph(pIJ699) (•) strain and the S. clavuligerus ORF15::aph(pIJ699-ORF15) (○) complemented strain. (Inset) Physical map of plasmid pIJ699-ORF15 used in the complementation studies. tsr and aph correspond to the thiostrepton and kanamycin resistance genes, respectively. Ter, terminator (from pIJ699). PtipA, tipA gene promoter. The restriction sites for HindIII (H), ApaI (A), SalI (S), and KpnI (K) are shown.
