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ORF1 of the S4 genome segment encodes the membrane fusion-inducing protein of BRV. Vero cells were transfected with pcDNA3 (a), pcDNA3-S4 (b), pcDNA3-S4 ORF1 (c), or pcDNA3-S4 ORF2 (d). Transfected monolayers were Wright-Giemsa stained 18 h posttransfection to detect the formation of multinucleated syncytia.
