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Linker hydrogen bonds as a function of spectrin length for the contraction-half of the NEMD cycle (slow trajectories, λ = 2000 ps; ξ = 0.010–0.040 shown as solid lines, ξ = 0.050–0.090 shown as dashed lines). The arrow indicates the direction of pulling (contraction). Error bars have been omitted for clarity, but are ∼±1 hydrogen bond.
