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A model for the evolution of Adh-Twain. (A) Chromosomal duplication of CG9010 in the ancestor of D. subobscura, D. madeirensis, and D. guanche. (B) Retrotransposition of an Adh mRNA into one of the copies of CG9010. Note that the 5′ regulatory regions of CG9010 are conserved. (C) Subsequent formation of a contiguous ORF spanning the remainder of CG9010 and the Adh retrosequence. The 3′ end of CG9010 is lost.
