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. 2005 Jun;170(2):841–857. doi: 10.1534/genetics.104.036384

TABLE 2.

Example of fitness matrix for diploid selection

00 01 11
00 1 1 + hs 1 + s
01 1 + hs (1 + hs)2 + ea×a (1 + hs)(1 + s) + 2ea×a + ea×d
11 1 + s (1 + hs)(1 + s) + 2ea×a + ea×d (1 + s)2 + 4ea×a + 4ea×d + ed×d

The entries correspond to the different genotypes at loci j and k; ea×a, ea×d, and ed×d correspond to additive × additive, additive × dominance, and dominance × dominance epistatic effects, respectively.