TABLE 1.

Maximum-likelihood two-locus model from parametric linkage analysis of the chromosome 16 F₂ intercross

	Position (cM)	Weight	Dominance	Variance explained (%)	Genetic effect of A/J allele
Proximal QTL	22.0	1.61	13.63	19	Overdominance; elevated PPI
Distal QTL	37.0	10.88	−9.82	23	Recessive; elevated PPI

Weight (additive component) and dominance (dominance component) of the QTL A/J allele (i.e., CSS-16 allele) effect in the following genetic model: Phenotype_i = mean + (Weight × Number of A/J alleles) + (Dominance × Het_i) + Noise, where Het_i = 1 if individual i is a heterozygote; otherwise Het_i = 0.