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Structure arrangement of the central four-helix bundle of TraM58–127. (A) Layers of a and d residues of the heptad repeats and water molecules. (B) Hydrogen bonding of the protonated Glu88 residues. (C) Hydrogen bonding of the central water molecules with the side-chain oxygen of Thr77 and the main-chain oxygen of Val74. (D) A stereo view of the 2Fo−Fc electron density map contoured at 1.5σ, showing residues Ile87 and Glu88 arranged around the four-fold axis of the tetramer.
