Mutation of methionine-43 to alanine in Snc1
yields a thermosensitive exocytic v-SNARE. (A)
SNC1ala43 encodes a thermosensitive v-SNARE.
Left panel: snc cells (SP1 background–JG8 and
W303 background–JG10) transformed with a control vector
(snc) or plasmids constitutively expressing
SNC1 or SNC1ala43 were
patched onto synthetic complete plates containing galactose. Cells were
grown for 3 d at 26°C, before replica plating onto medium
containing glucose to deplete SNC1. After 24 h,
patches were replica plated onto the following: galactose- (GAL) and
glucose-containing (GLU) plates at 26°C; prewarmed glucose-containing
plates at 35°C and 37°C; and amino acid-rich medium (YPD). Plates
were incubated for 30 h. Right panel: snc cells
(JG8) transformed with a control vector (snc), or
plasmids expressing SNC1 or
SNC1ala43, were grown to log phase on
galactose-containing medium and then were shifted to glucose-containing
medium for 24 h to deplete SNC1. Cells were seeded
in fresh glucose-containing medium at 26°C, and optical density
(measured at 600 nm) was monitored as a function of time. (B) Invertase
secretion is blocked in SNC1ala43 cells
after a shift to restrictive temperatures. Wild-type (SP1),
snc (JG8), and snc cells (JG8)
transformed with plasmids expressing SNC1 or
SNC1ala43 were grown to log phase on
galactose-containing medium, shifted to glucose medium for 24 h at
26°C and then derepressed for 2 h on low glucose (0.05%) medium
at either 26°C or 37°C. Both secreted (black) and nonsecreted
(gray) invertase activities were measured. (C)
SNC1ala43 cells accumulate secretory
vesicles at restrictive temperatures. snc cells (JG8)
expressing a control vector (snc), SNC1,
or SNC1ala43 were grown to log phase on
glucose-containing medium. Cells were maintained at 26°C, or were
shifted for 2 h to 37°C (indicated as 37°C), and were
processed for electron microscopy. Bar indicates 1 μm. (D)
Localization of Snc1-GFP and Snc1ala43-GFP in
sec6 cells. sec6–4 cells transformed
with plasmids expressing Snc1-GFP or Snc1ala43-GFP were
maintained at 26°C or were shifted for 2 h to 37°C, and then
were processed for confocal microscopy.