Expression of trophoblast marker genes in wild-type (WT) and RXRα−/−/RXRβ−/− (Xα/Xβ) placentas at E9.5. a–d and e–h are adjacent frozen sections stained with hematoxylin (a and e) or hybridized with antisense riboprobes for 4311, pl-1, and Mash-2 (b–d and f–h). C, chorionic plate; G, giant cell zone; L, labyrinthine zone; S, spongiotrophoblast; U, uterine epithelium.