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Cloverleaf structure of E. coli initiator Met-tRNAN. Nucleotides playing a major role in formylation are boxed by solid lines, whereas those playing a relatively minor role are boxed by dotted lines. RNase cleavages that are decreased in the presence of MTF are indicated with ⊝ whereas those that are increased are indicated with ⊕. 
, RNase V1 cleavage; 
, RNase T2 cleavage.
