Pollen and embryo development is arrested in ysl1yls3 mutants. A to B, Anthers stained for pollen viability. Viable pollen grains are stained magenta. A, Anther of ysl1ysl3 double mutant plant. B, Anther of Col-0 plant. C, Embryo arrest in seeds of ysl1ysl3. Seeds were imbibed, then dissected. Embryo from Col-0 (marked WT) surrounded by examples of embryos from ysl1ysl3 seeds.