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Proposed biosynthetic route for mannosylerythritol lipids. The first step of MEL biosynthesis is condensation of mannosyl and erythritol catalyzed by the glycosyltransferase Emt1. Mac1 and Mac2 are proposed to transfer short- and medium-chain fatty acids to positions R-2 and R-3. The last step, acetylation of deacetylated MEL at positions R-4 and R-6, is catalyzed by a single enzyme, Mat1.
