Functional complementation of the
molybdate-dependent murine cell line L929. L929 cells were transfected
with constructs expressing the full-length gephyrin cDNA clone P1
(+geph), the C-terminal truncated gephyrin
(+gephΔC) and Cnx1 (+cnx1), and pcDNA3
plasmid without insert (−) as a control. (A Upper) PCR
amplification of defined parts of the gephyrin and
cnx1 cDNA sequences in different transgenic cell lines.
Amplifications were performed by using genomic DNA of two independent
pools of transformants (P1, P2), of untransfected L929 cells in the
negative control (n.c.), and plasmid DNA of the different expression
constructs as positive control (p.c.). (Lower) Analysis
of transgene expression by RNase protection assays in cells stably
transfected. In the case of the L929 cells stably transfected with the
gephyrin expression constructs (geph and
gephΔC), an increase of the gephyrin mRNA level in
comparison to the untransfected L929 cells can be seen, demonstrating
the expression of the transgene. (B) Moco content of
transfected L929 cells. The columns show Moco activity obtained from
multiple nit-1 complementation assays of NR6P cells
(black bar), L929 cells (stippled bars) grown in the presence (+Mo) or
absence (−Mo) of 1 mM sodium molybdate and pools of at least 100
stable transformants (gray bars). 100% of wild-type Moco content
represent a NADPH–nitrate reductase activity of 51 ± 4.5
nmol/mg protein⋅h in the nit-1 complementation
assay. (C) Antisense expression of gephyrin cDNA in murine
cells. For gephyrin depletion, cells of the murine wildtype cell line
NR6P were transfected with the pcDNA3 gephyrin antisense expression
construct and as a negative control with pcDNA3 without insert
(indicated by −), and stable transformants were selected. The colums
show Moco contents of L929 cells, untransfected NR6P cells, NR6P cells
transfected with control vector (NR6P−), and cell clones derived from
the transfection with the antisense construct (NR6Pgeph
antisense clone A and clone B) grown on normal medium (−Mo, white and
gray bars) and on medium supplemented with 1 mM sodium molybdate for
12 h (+Mo, stippled and black bars). Values are expressed in % of the Moco content of the untransformed
NR6P wild-type cells (51 ± 4.5 nmol/mg protein⋅h).