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A heterologous plaque assay was performed to illustrate the spreading of L. monocytogenes from infected P388.D1 macrophages into a HepG2 cell layer. Plaques were imaged 3 days after infection. Whereas the wild-type (wt) bacteria produced plaques, L. monocytogenes Δiap was unable to spread from the macrophages to the hepatocytes and did not show plaque formation.
