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Various substrates that were also incubated with the 20,000g supernatant and NADPH. A mixture of α-costic acid (20a) and γ-costic acid (20b) or elema-1,3,11(13)-trien-12-oic acid (21) may serve as substrate analogs of germacrene acid (14). Parthenolide (22) and dehydrocostus lactone (23) serve as model compounds for conversions that might occur during chicory sesquiterpene lactone biosynthesis after formation of (+)-costunolide.
