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. 2005 Nov 1;272(1581):2561–2569. doi: 10.1098/rspb.2005.3356

Table 1.

Review on studies on shifts in phenology where a yardstick to assess whether these shifts are sufficient can be identified.

species phenological event yardstick response reference
D. pulicaria (fresh water zooplankton) abundance diatom (A. formosa) bloom too little Winder & Schindler (2004)
K. cochlearis (fresh water zooplankton) abundance diatom (A. formosa) bloom sufficient Winder & Schindler (2004)
M. balthica (a marine intertidal bivalve) spawning phytoplankton bloom and settlement of juvenile shrimps (Cragon cragon) unclear Philippart et al. (2003)
winter moth (O. brumata) egg hatching oak (Q. robur) bud burst too much Visser & Holleman (2001)
orange tip butterfly (A. cardamines) first appearance garlic mustard (A. petiolata) flowering date sufficient (?) Sparks & Yates (1997), Harrington et al. (1999)
red admiral (V. atalanta) migration–return date stinging nettle (U. dioica) flowering too much Sparks et al. (2005)
great tit (P. major) laying date caterpillar biomass peak too little Visser et al. (1998), Visser et al. (in press b)
great tit (P. major) laying date caterpillar biomass peak sufficient Cresswell & McCleery (2003)
golden plover (P. apricaria) laying date insect abundance peak unknown Pearce-Higgins et al. (2005)
black-legged kittiwake (R. tridactyla), common guillemot (U. aalge), European shag (P. aristotelis) laying date sandeel (Ammadytes marinus) abundance peak unknown Rindorf et al. (2000), Frederiksen et al. (2004)
puffin (F. artica) laying date herring (C. harengus) abundance peak unknown Durant et al. (2003)
wood warblers (Parulidae) migration eastern spruce budworm (C. fumiferana) density peak too little Strode (2003)
pied flycatcher (F. hypoleuca) migration–arrival date caterpillar biomass peak too little Both & Visser (2001)
honey-buzzard (P. apivorus) migration–arrival date wasp (Vespula sp.) abundance peak too little R. Bijlsma (unpublished data)
American robin (T. migratorius) migration–arrival date snow melt too much Inouye et al. (2000)

Responses are classified as either too strong (the shift is larger than the shift in the yardstick), sufficient or too weak. In some cases, the shift in the yardstick is not reported and these are classified as unknown. In the Macoma example, it is unclear whether the shift is sufficient, see text. For an example on a marine pelagic community (Edwards & Richardson 2004), see text.