TABLE 1.

Definition of parameters and notation used in text

Parameter or term	Definition
AiAj	Individual with maternally inherited Ai allele and paternally inherited Aj allele
Aijkl	AiAj offspring with an AkAl mother
Gijkl	Genotypic value of Aijkl
frijkl	Frequency of Aijkl in population
gdijkl	Genotypic deviation for Aijkl, the difference between the genotypic value (Gijkl) and the population mean
	Breeding value of female AiAj genotype; breeding value of male AiAj genotype
	Dominance deviation for female Aijkl; dominance deviation for male Aijkl
	Total genetic variance of population
;	Additive genetic variance for females; additive genetic variance for males
;	Dominance genetic variance for females; dominance genetic variance for males
;	Covariance between breeding values (additive effects) and dominance deviations for females and for males
;	Covariance between offspring and mother (female parent) genotypic values; covariance between offspring and father (male parent) genotypic values
	Covariance between full-sib genotypic values
;	Covariance between genotypic values of half-sibs sharing a female parent and of those sharing a male parent
;	Additive effect of inheriting an Ai allele from the mother; additive effect of inheriting an Aj allele from the father
;	Additive effect of having a mother who received an Ak allele from her mother; additive effect of having a mother who received an Al allele from her father
	Average genotypic value of AiAj genotype
	Average genetic value of individuals with an AkAl mother
; ;	Dominance effect of AiAj genotype; dominance effect for individual with AkAl mother; combined offspring–mother genotype dominance effect
;	Offspring genotype additive genetic variation; maternal genotype additive genetic variation
; ;	Offspring genotype dominance genetic variance; maternal genotype dominance genetic variation; combined offspring–mother genotype dominance genetic variation
;	Covariance between additive and additive effects; covariance between additive and dominance effects