TABLE 2.

Expressions of quadrigenic disequilibrium D_AB in terms of genotypic configuration frequencies, allele frequencies, and lower-order disequilibrium coefficients

Frequency	1		DA	DB	Dab	Da/b	DAb	DaB
		−1			−2pApB	−2pApB	−2pB	−2pA
		−1	pBpb		−pApB + pApb	−pApB + pApb	−pB + pb	−2pA
		−1			2pApb	−2pApb	−2pb	−2pA
		−1		pApa	−pApB + papB	−pApB + papB	−2pB	−pA + pa
	−pApapBpb	−1	pBpb	pApa	−pApB − papb	pApb + papB	−pB + pb	−pA + pa
	−pApapBpb	−1	pBpb	pApa	pApb + papB	−pApB − papb	−pB + pb	−pA + pa
		−1		pApa	pApb − papb	pApb − papb	2pb	−pA + pa
		−1			2papB	2papB	−2pB	2pa
		−1	pBpb		papB − papb	papB − papb	−pB + pb	2pa
		−1			−2papb	−2papb	2pb	2pa