TABLE 3.

Expressions of composite quadrigenic disequilibrium Δ_AB in terms of genotypic and allele frequencies and lower-order disequilibrium coefficients

Frequency	1		DA	DB	Δab	DAb	DaB
		−1			−2pApB	−2pB	−2pA
		−1	pBpb		−pApB + pApb	−pB + pb	−2pA
		−1			2pApb	2pb	−2pA
		−1		pApa	−pApB + papB	−2pB	−pA + pa
a	−pApapBpb	−1	pBpb	pApa		−pB + pb	−pA + pa
		−1		pApa	pApb − papb	2pb	−pA + pa
		−1			2papB	−2pB	2pa
		−1	pBpb		papB − papb	−pB + pb	2pa
		−1			−2papb	2pb	2pa

^a.