Correlation between linkage disequilibrium and physical distance in four scorpion species. (Estimates of R2 and |D′| were carried out using the computer package ‘LDhat’. p values were calculated from a one-sided Mantel test with 10 000 permutations. Genes were tested in the order 16S-5 kb-coxI using the shorter distance. Estimates do not change remarkably if they are tested using the longer connection of approximately 10 kb. ***p values <0.000 1, **p values >0.000 1 and <0.01, and *p values >0.01 and <0.05.)