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The block design used to measure inbreeding depression. Each block is created by crossing beetles from two unrelated families, creating two outbred matings (reciprocal crosses between the two families) and two inbred matings (crosses between full-sibs within each family). Outbreds and inbreds within each block thus have, on average, the same set of alleles but differ in degree of homozygosity due to the mating treatment.
