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Phylogeny and stratigraphic record of sea turtles, including Bouliachelys (see electronic supplementary material). Living lineages in black lines, extinct lineages in grey. Boxed numbers at selected nodes refer to synapomorphies discussed in the main text. Other numbers refer to bootstrap/Bremer support. The deep nesting of the ancient Santanachelys suggests the existence of at least five lineages in the Aptian (depending on how polytomies are resolved).
